Morphological phylogeny of Megachilini and the evolution of leaf-cutter behavior in bees (Hymenoptera: Megachilidae) Author Gonzalez, Victor H. Undergraduate Biology Program, Haworth Hall, 1200 Sunnyside Avenue, University of Kansas, Lawrence, Kansas 66045, USA (victorgonzab @ gmail. com). & Department of Ecology & Evolutionary Biology, Haworth Hall, 1200 Sunnyside Avenue, University of Kansas, Lawrence, Kansas 66045, USA (gtgustafson @ gmail. com). Author Gustafson, Grey T. Department of Ecology & Evolutionary Biology, Haworth Hall, 1200 Sunnyside Avenue, University of Kansas, Lawrence, Kansas 66045, USA (gtgustafson @ gmail. com). & Division of Entomology, Natural History Museum, 1501 Crestline Drive - Suite 140, University of Kansas, Lawrence, Kansas 66045 - 4415, USA (msengel @ ku. edu). Author Engel, Michael S. Department of Ecology & Evolutionary Biology, Haworth Hall, 1200 Sunnyside Avenue, University of Kansas, Lawrence, Kansas 66045, USA (gtgustafson @ gmail. com). & Division of Entomology, Natural History Museum, 1501 Crestline Drive - Suite 140, University of Kansas, Lawrence, Kansas 66045 - 4415, USA (msengel @ ku. edu). & Division of Invertebrate Zoology, American Museum of Natural History, Central Park West at 79 Street, New York, New York 10024 - 5192, USA. text Journal of Melittology 2019 2019-07-03 2019 85 1 123 http://dx.doi.org/10.17161/jom.v0i85.11541 journal article 10.17161/jom.v0i85.11541 2325-4467 13228910 46F71985-5AF2-4AF8-AD53-1E9070547021 Ochreriadini Gonzalez & Engel , new tribe ZooBank: urn:lsid:zoobank.org:act: D70C4AF0-CE03-4A89-9881-AA8ABAE29225 ( Fig. 15 ) TYPE GENUS : Ochreriades Mavromoustakis, 1956 . DIAGNOSIS: This tribe is readily separated from all other tribes of Megachilinae by the following combination of features: body elongate and with yellow or ivory integumental markings; pronotum distinctly elevated and surrounding mesoscutum anteriorly; mouthparts elongate, reaching tip of metasoma. DESCRIPTION: ♀ : Clypeus not overhanging labral base; labrum not elongate, margin without fringe or apical tuft of setae; maxillary palpus trimerous (three palpomeres); metepisternum with dorsal carina or lamella; pronotum enlarged and surrounding mesoscutum anteriorly, practically eliminating omaular surface of mesepisternum and anterior surface of mesoscutum; mesoscutellum flat, on same plane with metanonum and propodeal base, as seen in profile; T6 without wide apical hyaline rim; S6 without lateral or apical projection. ♂ : Metasoma with six sterna visible; S2 and S3, each with disc swollen; S4 with dense pubescence on disc; S5 not emarginate, with branched or simple discal setae; T7 exposed, inferiorly directed . COMMENTS: This tribe contains a single genus, Ochreriades , which consists of two species. Ochreriades fasciatus (Friese) occurs in the Middle East whereas O . rozeni Griswold occurs in Namibia , Africa ( Griswold, 1994 ; Ascher & Pickering, 2018).