Macrodactylini (Coleoptera, Scarabaeidae, Melolonthinae): primary types of type species and taxonomic changes to the generic classification
Author
Fuhrmann, Juares
Author
Vaz-De-Mello, Fernando Z.
text
European Journal of Taxonomy
2017
2017-09-13
350
1
71
journal article
22059
10.5852/ejt.2017.350
52e4ae01-f51f-4c4a-8a14-9fb5b17788d6
2118-9773
3832682
8D14DBDE-AD13-445B-B2D0-221F19FC7C37
Paulosawaya
Martínez & d’Andretta, 1956
, revalidated
Amphicrania
Burmeister, 1855: 26
(
non Amphicania
Dejean, 1833, a synonym of
Liogenys
,
Diplotaxini
),
syn. nov
.
Type
species:
Amphicrania ursina
Burmeister, 1855
(designation:
Evans 2003: 249
); junior synonymy of
Paulosawaya ursina
(
Blanchard, 1850
)
comb. nov.
(original combination:
Clavipalpus ursinus
Blanchard, 1850
).
Pseudoleuretra
Martínez & d’Andretta, 1956: 351
,
syn. nov
.
Type
species:
Pseudoleuretra bokermanni
Martínez & d’Andretta, 1956
(original designation).
Amphicrania
–
Gemminger & Harold 1869: 1143
(synonymy with
Clavipalpus
).
Paulosawaya
–
Smith 2008: 22
(synonymy with
Clavipalpus
).
Type
species
Paulosawaya ornatissima
Martínez & d’Andretta, 1956
(original designation).
Diagnosis
Clypeal ventral area strongly widened; maxillary palpomere IV elongate (
Fig. 6D
); pronotal anterior and posterior margins not beaded; prosternum anteriorly concave (similar to
Fig. 12K
); protibia with 2–3 external teeth and without spur; elytral striae indistinct, posterior and posterointernal margins not beaded; meso- and metatibiae each with a transverse carina; metatarsomere V without spine-like setae; abdomen with intersegmental membrane VII–VIII concealed.
Remarks
Katovich (2008)
redescribed
Clavipalpus
based on
syntypes
of
C
.
ursinus
Blanchard, 1850
and the
holotype
of
C
.
aequatorialis
Moser, 1918
. Thereafter,
Smith (2008)
proposed the synonymy of
Paulosawaya
with
Clavipalpus
based on the study of
P
.
ornatissima
(type species of
Paulosawaya
) and
Clavipalpus
spp.
Clavipalpus ursinus
,
C
.
aequatorialis
and
P
.
ornatissima
evidently belong to the same genus as stated by
Smith (2008)
, but the problem is that the type material of the type species of the genus
Clavipalpus
(
C
.
dejeani
) was not examined by these studies (
Katovich 2008
;
Smith 2008
).
The present study designates a
lectotype
for
Clavipalpus dejeani
and
Amphicrania ursinus
, revalidates
Paulosawaya
(type species:
P
.
ornatissima
,
Fig. 22
A–B), and proposes the synonymy of
Amphicrania
Burmeister
and
Pseudoleuretra
(type species:
P
.
bokermanni
,
Fig. 22
C–D) with
Paulosawaya
.
Amphicrania
Burmeister
is a senior synonym of
Paulosawaya
, but it is a junior homonym of
Amphicrania
Dejean
(a synonym of
Liogenys
according to
Erichson 1848
) and may not be used as a valid name (
ICZN 1999
: Articles 23.3.5. and 52.2).
Paulosawaya
now comprises 17 species and one subspecies:
P
.
aequatorialis
(
Moser, 1918
)
comb. nov.
,
P
.
antisanae
(Bates, 1891)
comb. nov.
,
P
.
basalis
(
Moser, 1918
)
comb. nov.
,
P
.
blanchardi
(Kirsch, 1885)
comb. nov.
,
P
.
bokermanni
(
Martínez & d’Andretta, 1956
)
comb. nov.
,
P
.
castanea
(
Moser, 1924
)
comb. nov.
,
P
.
cochleata
(
Burmeister, 1855
)
comb. nov.
,
P
.
hirsuta
(Kirsch, 1885)
comb. nov.
,
P
.
ornatissima
Martínez & d’Andretta, 1956
,
P
.
peruana
(
Moser, 1918
)
comb. nov.
,
P
.
rimbachi
(
Moser, 1924
)
comb. nov.
,
P
.
sinuatus
(Kirsch, 1885)
comb. nov.
,
P
.
spadicea
(
Burmeister, 1855
)
comb. nov.
,
P
.
ursina
(
Blanchard, 1850
)
comb. nov.
,
P
.
variolosa
(
Burmeister, 1855
)
comb. nov.
,
P
.
vestita
(
Moser, 1924
)
comb. nov.
,
P
.
whymperi chimborazonus
(Bates, 1891)
comb. nov.
and
P
.
whymperi whymperi
(Bates, 1891)
comb. nov.
The present study revalidates
Paulosawaya
, formerly a synonym of
Clavipalpus
, based on the following characters (opposition to
Clavipalpus
): clypeus broad and semicircular or truncated (narrow and trapezoid); clypeal posterior angle not extended over the canthus (partially covering the canthus); maxillary palpus long (
Fig. 6D
) (short,
Fig. 6B
), palpomere IV distinctly enlongate (
Fig. 6D
) (enlarged,
Fig. 6B
); meso- and metatibiae each with a raised transverse carina (carina distinct by large punctation).
Paulosawaya
usually includes large specimens (over
12 mm
), that are reddish-brown, with long setae and occuring in northern
Brazil
,
Guyana
,
Peru
,
Ecuador
and
Colombia
, and
Clavipalpus
includes medium-sized specimens (about
9 mm
), yellowish-brown, with short setae and occuring in
Brazil
.