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<mods:title id="3044396195C174C50340E36993941838">On the types species of the New World dung beetle genus Canthidium Erichson, 1847 (Coleoptera: Scarabaeidae: Scarabaeinae), with an annotated checklist of species</mods:title>
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<taxonomicName id="1381E691FFCCFFEEFF2D52401D6CF98E" ID-CoL="QNB6" authority="Howden &amp; Young, 1981" authorityName="Howden &amp; Young" authorityYear="1981" box="[151,886,1627,1653]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="464" phylum="Arthropoda" rank="species" species="tenebrosum" subGenus="Canthidium">
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<emphasis id="E6F54100FFCCFFEEFF2D52401F38F98F" bold="true" box="[151,290,1627,1652]" italics="true" pageId="13" pageNumber="464">Canthidium</emphasis>
(
<emphasis id="E6F54100FFCCFFEEFE8852401FA7F98F" bold="true" box="[306,445,1627,1652]" italics="true" pageId="13" pageNumber="464">Canthidium</emphasis>
) 
<emphasis id="E6F54100FFCCFFEEFE7652401C4EF98F" bold="true" box="[460,596,1627,1652]" italics="true" pageId="13" pageNumber="464">tenebrosum</emphasis>
<bibRefCitation id="B010E0E3FFCCFFEEFDE152471D6CF98E" box="[603,886,1628,1653]" pageId="13" pageNumber="464" refId="ref27924" refString="Howden, H. F. &amp; Young, O. P. (1981) Panamanian Scarabaeinae: taxonomy, distribution, and habits (Coleoptera, Scarabaeidae). Contributions of the American Entomological Institute, 18, 1 - 204." type="journal article">Howden &amp; Young, 1981</bibRefCitation>
</emphasis>
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<emphasis id="E6F54100FFCCFFEEFF7D529B1FC2F963" box="[199,472,1664,1688]" italics="true" pageId="13" pageNumber="464">Canthidium tenebrosum</emphasis>
<treatmentCitation id="5520BB03FFCCFFEEFE5852641D34F963" author="" box="[482,814,1663,1688]" page="82" pageId="13" pageNumber="464">
<bibRefCitation id="B010E0E3FFCCFFEEFE5852641D1EF963" box="[482,772,1663,1688]" pageId="13" pageNumber="464" refId="ref27924" refString="Howden, H. F. &amp; Young, O. P. (1981) Panamanian Scarabaeinae: taxonomy, distribution, and habits (Coleoptera, Scarabaeidae). Contributions of the American Entomological Institute, 18, 1 - 204." type="journal article">Howden &amp; Young, 1981</bibRefCitation>
: 82
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</taxonomicName>
[here transferred to the subgenus 
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<emphasis id="E6F54100FFCCFFEEFB78529B1B5CF963" box="[1218,1350,1664,1688]" italics="true" pageId="13" pageNumber="464">Canthidium</emphasis>
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]. 
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locality:
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<collectingRegion id="164553F0FFCCFFEEFEBC52BE1F79F947" box="[262,355,1701,1724]" country="Panama" name="Panama" pageId="13" pageNumber="464">
<collectingCountry id="AC96DD82FFCCFFEEFEBC52BE1F79F947" box="[262,355,1701,1724]" name="Panama" pageId="13" pageNumber="464">Panama</collectingCountry>
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: 
<collectingRegion id="164553F0FFCCFFEEFECF52BF1FCFF946" box="[373,469,1700,1725]" country="Panama" name="Chiriqui" pageId="13" pageNumber="464">Chiriquí</collectingRegion>
, 
<quantity id="137930F7FFCCFFEEFE5252BF1C33F946" box="[488,553,1700,1725]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="464" unit="km" value="2.0">2 km</quantity>
west of Cerro Punta, “Baldwin Forest”. 
<emphasis id="E6F54100FFCCFFEEFB9D52BF1AC5F946" bold="true" box="[1063,1247,1700,1725]" pageId="13" pageNumber="464">
<typeStatus id="0B3A23B0FFCCFFEEFB9D52BF1A78F946" box="[1063,1122,1700,1725]" pageId="13" pageNumber="464">Type</typeStatus>
material:
</emphasis>
<typeStatus id="0B3A23B0FFCCFFEEFB5452BF1B4CF946" box="[1262,1366,1700,1725]" pageId="13" pageNumber="464" type="holotype">Holotype</typeStatus>
male (CMNC—ex Howden collection); see full list of 
<typeStatus id="0B3A23B0FFCCFFEEFD7D52D31D29F91B" box="[711,819,1736,1760]" pageId="13" pageNumber="464" type="paratype">paratypes</typeStatus>
in 
<treatmentCitation id="5520BB03FFCCFFEEFCE052DC1AB1F91B" author="" box="[858,1195,1735,1760]" page="83" pageId="13" pageNumber="464">
<bibRefCitation id="B010E0E3FFCCFFEEFCE052DC1AB1F91B" box="[858,1195,1735,1760]" pageId="13" pageNumber="464" refId="ref27924" refString="Howden, H. F. &amp; Young, O. P. (1981) Panamanian Scarabaeinae: taxonomy, distribution, and habits (Coleoptera, Scarabaeidae). Contributions of the American Entomological Institute, 18, 1 - 204." type="journal article">Howden &amp; Young (1981: 83)</bibRefCitation>
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. 
<emphasis id="E6F54100FFCCFFEEFB0252DC1B4EF91B" bold="true" box="[1208,1364,1735,1760]" pageId="13" pageNumber="464">Distribution:</emphasis>
<collectingCountry id="AC96DD82FFCCFFEEFAE652D31ED1F8FE" name="Costa Rica" pageId="13" pageNumber="464">Costa Rica</collectingCountry>
and 
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<collectingCountry id="AC96DD82FFCCFFEEFEBB52F61F40F8FF" box="[257,346,1773,1796]" name="Panama" pageId="13" pageNumber="464">Panama</collectingCountry>
</collectingRegion>
(Solís &amp; Kohlmann 2004).
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