Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae) Author Johnson, Andrew J. SchoolofForestResourcesandConservation, UniversityofFlorida, Gainesville, FL 32611, andrewjavanjohnson@gmail.com Author Hulcr, Jiri SchoolofForestResourcesandConservation, UniversityofFlorida, Gainesville, FL 32611, Author Knížek, Miloš Department of Forest Protection Service, Forestry and Game Management Research Institute, Strnady, Jíloviště, Praha 5, Zbraslav CZ- 15600, Czechia, Author Atkinson, Thomas H. Texas Natural History Collections, Integrative Biology, Universityof Texasat Austin, Austin, TX 78712, Author Mandelshtam, Michail Yu. Department of Forest Protection, Wood Science and Game Management, Saint-Petersburg State Forest Technical University named after S. M. Kirov, Institutskii per., 5,194021 Saint-Petersburg, Russia, Author Smith, Sarah M. Department of Entomology, Michigan State University, East Lansing, MI 48824, Author Cognato, Anthony I. Department of Entomology, Michigan State University, East Lansing, MI 48824, Author Park, Sangwook Research Institute of Forest Insect Diversity, Namyangju 12113, South Korea, Author Li, You SchoolofForestResourcesandConservation, UniversityofFlorida, Gainesville, FL 32611, Author Jordal, Bjarte H. UniversityMuseum ofBergen, University of Bergen, P. O. 7800,5020 Bergen, and text Insect Systematics and Diversity 2020 2020-05-01 4 3 1 1 81 http://dx.doi.org/10.1093/isd/ixaa002 journal article 10.1093/isd/ixaa002 2399-3421 Cryphalus Erichson, 1836: 61 ( Figs. 10–12 ) Synonymy = Pseudocryphalus Ferrari, 1869: 252 . = Taenioglyptes Bedel, 1888: 398 . = Cryptarthrum Blandford, 1896a: 200 . = Allarthrum Hagedorn, 1912c: 355 . = Dacryphalus Hopkins, 1915: 42 syn. nov. = Ericryphalus Hopkins, 1915: 38 . = Hypocryphalus Hopkins, 1915: 41 syn. nov. = Margadillius Hopkins, 1915: 37 syn. nov. = Piperius Hopkins, 1915: 39 . = Toenioglyptes Balachowsky, 1949: 205 (unavailable name). = Ernocryphalus Murayama, 1958: 934 . = Acryphalus Tsai and Li, 1963: 604 (unavailable name). = Gugocryphelus Tsai and Li, 1963: 603 (unavailable name). = Jugocryphalus Tsai and Li, 1963: 604 . Type of genus Bostrichus asperatus Gyllenhal, 1813 . Diagnosis Cryphalus can be distinguished from similar genera by the combination of emarginated eyes, sutures on the antennae clearly visible but without a septum, by the weakly bilobed third tarsal segments, the proventriculus with a large apical plate, and the aedeagus that has a tegmen sclerotized completing a ring, usually with two tegminal apodemes.