<document id="55060DE7CCDD7FAD0CCA16B78107A866" ID-CLB-Dataset="8980" ID-DOI="10.11646/zootaxa.4820.3.2" ID-GBIF-Dataset="8f91a52d-de34-47ed-80ac-9dc7c1bf915c" ID-ISSN="1175-5326" ID-Zenodo-Dep="4398015" ID-ZooBank="560370FE-B271-48D2-A720-DB9034CE308E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609211755247" checkinUser="plazi" docAuthor="Kluge, Nikita J." docDate="2020" docId="960C8792160D753056E3FCB7B4E3BA79" docLanguage="en" docName="zootaxa.4820.3.2.pdf" docOrigin="Zootaxa 4820 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Demoulinia Gillies 1990" docType="treatment" docVersion="8" lastPageNumber="439" masterDocId="6A35FFEA160C75315674FFF5B178BE78" masterDocTitle="Demoulinia Gillies 1990 and two unnamed genera of the plesiomorphon Protopatellata (Ephemeroptera, Baetidae) from South Africa" masterLastPageNumber="464" masterPageNumber="438" pageNumber="439" updateTime="1698848356202" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="388AB6991C4ED1EB422C6D3B96021B1F">Demoulinia Gillies 1990 and two unnamed genera of the plesiomorphon Protopatellata (Ephemeroptera, Baetidae) from South Africa</mods:title>
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<taxonomicName id="D9A54D07160D753056E3FCB7B0B8BD24" ID-CoL="42WK" authority="Gillies 1990" authorityName="Gillies" authorityYear="1990" box="[151,448,834,861]" class="Insecta" family="Baetidae" genus="Demoulinia" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="439" phylum="Arthropoda" rank="genus">
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<emphasis id="2CD1EA96160D753056E3FCB7B05EBD25" bold="true" box="[151,294,834,861]" italics="true" pageId="1" pageNumber="439">Demoulinia</emphasis>
<bibRefCitation id="7A344B75160D7530575AFCB7B0B8BD24" author="Gillies, M. T." box="[302,448,834,860]" pageId="1" pageNumber="439" pagination="97 - 128" refId="ref10174" refString="Gillies, M. T. (1990) A revision of the African species of Centroptilum Eaton (Baetidae, Ephemeroptera). Aquatic Insects, 12 (2), 97 - 128. https: // doi. org / 10.1080 / 01650429009361395" type="journal article" year="1990">Gillies 1990</bibRefCitation>
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(
<figureCitation id="869E2A01160D753056EBFC92B068BDF9" box="[159,272,871,897]" captionStart-0="FIGURES 1–5" captionStart-1="FIGURES 6–12" captionStart-10="FIGURES 62–66" captionStart-11="FIGURES 67–70" captionStart-12="FIGURE 71" captionStart-2="FIGURES 13–20" captionStart-3="FIGURES 21–25" captionStart-4="FIGURES 26–31" captionStart-5="FIGURES 32–38" captionStart-6="FIGURES 39–44" captionStart-7="FIGURES 45–51" captionStart-8="FIGURES 52–56" captionStart-9="FIGURES 57–61" captionStartId-0="5.[151,264,1889,1913]" captionStartId-1="6.[151,264,1860,1884]" captionStartId-10="17.[151,264,1702,1726]" captionStartId-11="18.[151,264,1865,1889]" captionStartId-12="19.[151,250,1495,1519]" captionStartId-2="8.[151,264,1867,1891]" captionStartId-3="9.[151,264,1790,1814]" captionStartId-4="10.[151,264,1769,1793]" captionStartId-5="11.[151,264,1921,1945]" captionStartId-6="12.[151,264,1894,1918]" captionStartId-7="13.[151,264,1996,2020]" captionStartId-8="14.[151,264,1747,1771]" captionStartId-9="16.[151,264,1374,1398]" captionTargetBox-0="[151,1434,181,1866]" captionTargetBox-1="[151,1436,181,1836]" captionTargetBox-10="[151,1429,193,1667]" captionTargetBox-11="[151,1435,181,1841]" captionTargetBox-12="[151,1436,181,1471]" captionTargetBox-2="[151,1436,292,1844]" captionTargetBox-3="[151,1436,181,1767]" captionTargetBox-4="[151,1436,181,1745]" captionTargetBox-5="[151,1436,181,1898]" captionTargetBox-6="[151,1436,181,1871]" captionTargetBox-7="[169,1415,424,1960]" captionTargetBox-8="[151,1436,181,1723]" captionTargetBox-9="[151,1436,310,1351]" captionTargetId-0="figure-32@5.[151,1436,181,1866]" captionTargetId-1="figure-16@6.[151,1436,181,1836]" captionTargetId-10="figure-32@17.[151,1435,181,1678]" captionTargetId-11="figure-16@18.[151,1435,181,1841]" captionTargetId-12="figure-32@19.[151,1436,181,1471]" captionTargetId-2="figure-56@8.[151,1436,273,1844]" captionTargetId-3="figure-32@9.[151,1436,181,1767]" captionTargetId-4="figure-16@10.[151,1436,181,1745]" captionTargetId-5="figure-32@11.[151,1436,181,1898]" captionTargetId-6="figure-16@12.[151,1436,181,1871]" captionTargetId-7="figure-127@13.[161,1425,417,1972]" captionTargetId-8="figure-16@14.[151,1436,181,1723]" captionTargetId-9="figure-80@16.[151,1436,309,1351]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-10="17" captionTargetPageId-11="18" captionTargetPageId-12="19" captionTargetPageId-2="8" captionTargetPageId-3="9" captionTargetPageId-4="10" captionTargetPageId-5="11" captionTargetPageId-6="12" captionTargetPageId-7="13" captionTargetPageId-8="14" captionTargetPageId-9="16" captionText-0="FIGURES 1–5. Larva of Demoulinia crassi. 1, Female larva of last instar; 2–3, exuviae of male larvae: 2, left half of prothorax and mesothorax; 3, abdomen; 4, abdomen of male subimago extracted from mature larva (to show hypodermal larval coloration); 5, tergalius III." captionText-1="FIGURES 6–12. Labrum of Demoulinia spp., dorsal view. 6–7, D. insularis (holotype). 8–12, D. crassi." captionText-10="FIGURES 62–66. Male genitalia of Demoulinia crassi. 62, penis, ventral view (left muscles not shown); 63, genitalia, ventral view (penis dotted; left gonovectal muscle and right styligeral muscle not shown); 64, penis (dotted), posterior view (left muscle not shown); 65, subimaginal gonostyli crumpled under larval cuticle; 66, exuviae subimaginal of gonostyli (gd, gonoduct; gs1, gs2, gs3, segments of gonostylus; m.gv, gonovectal-sternal muscle; m.g-t, gonovectal-tergal muscle; m.pp, paces of attachment of muscles going to paraprocts; m.sg, sterno-styligeral muscle; usg, unistyliger)." captionText-11="FIGURES 67–70. Females of Demoulinia crassi. 67–68, female imago; 69–70, subimaginal exuviae of tibia and tarsus: 69, middle leg; 70, fore leg." captionText-12="FIGURE 71. Egg of Demoulinia crassi." captionText-2="FIGURES 13–20. Mandibles of Demoulinia crassi. 13, 16, 19, left mandible; 14, 17, 20, right mandible; 15, apex of right mandible, proximal view (inc, incisor; kd, kinetodontium; prs, prostheca)." captionText-3="FIGURES 21–25. Demoulinia spp. 21–23, D. crassi; 24–25, D. insularis (holotype); 21, 22, 24, maxilla (21, with focus on ventral side; 22, with focus on dorsal side; d1, d2, d3, dentisetae; p1, p2, p3, segments of maxillary palp); 23, 25, hypopharynx and superlingua." captionText-4="FIGURES 26–31. Labium of Demoulinia crassi. 26, cross section of glossae and paraglossae; 27, labium and maxilla; 28–31, exuviae of glossae, paraglossae and labial palp: 28–29, with focus on ventral side; 30–31, with focus on dorsal side (gl:dl, dorsolateral row of stout setae on glossa; gl:m, median row of stout setae on glossa; gl:vm, setae corresponding to ventro-median row of glossa; pg:dm, dorso-median row of long setae on paraglossa; pg:la, latero-apical long setae of paraglossa; pg:vm, long setae corresponding to ventro-median row of paraglossa)." captionText-5="FIGURES 32–38. Larval legs of Demoulinia spp. 32–37, D. crassi: 32, exuviae of hind leg; 33, inner side of fore tarsus; 34, femoral-tibial of hind leg, focus on anterior side; 35–37, claws. 38, D. insularis (holotype), fragment of claw with subimaginal claws inside (arrow shows row of vestigial denticles on inner side)." captionText-6="FIGURES 39–44. Exuviae of larval abdomen of Demoulinia spp. 39–43, D. crassi: 39, lateral and posterior area of abdominal tergum VII; 40, anterior-lateral area of abdominal sternum VI; 41, caudalii; 42, fragment of cercus; 43, paraproct. 44, D. insularis (holotype), median margin of paraproct (scale bar refers to Figs 39, 40, 42, 43)." captionText-7="FIGURES 45–51. Demoulinia crassi, tergalii I–VII." captionText-8="FIGURES 52–56. Demoulinia crassi. 52–53, male imago; 54–56, subimaginal exuviae: 54, left half of mesonotum; 55, abdomen; 56, fragment of pleuron with postsubalar sclerite." captionText-9="FIGURES 57–61. Male genitalia of Demoulinia spp. 57, D. insularis (paratype of D. assimilis), subimaginal gonostyli under larval cuticle. 58–61, D. crassi: 58, subimaginal gonostyli extracted from larva; 59, imaginal genitalia; 60–61, apex of imaginal gonostylus." figureDoi-0="http://doi.org/10.5281/zenodo.4398017" figureDoi-1="http://doi.org/10.5281/zenodo.4398019" figureDoi-10="http://doi.org/10.5281/zenodo.4398039" figureDoi-11="http://doi.org/10.5281/zenodo.4398041" figureDoi-12="http://doi.org/10.5281/zenodo.4398043" figureDoi-2="http://doi.org/10.5281/zenodo.4398023" figureDoi-3="http://doi.org/10.5281/zenodo.4398025" figureDoi-4="http://doi.org/10.5281/zenodo.4398027" figureDoi-5="http://doi.org/10.5281/zenodo.4398029" figureDoi-6="http://doi.org/10.5281/zenodo.4398031" figureDoi-7="http://doi.org/10.5281/zenodo.4398033" figureDoi-8="http://doi.org/10.5281/zenodo.4398035" figureDoi-9="http://doi.org/10.5281/zenodo.4398037" httpUri-0="https://zenodo.org/record/4398017/files/figure.png" httpUri-1="https://zenodo.org/record/4398019/files/figure.png" httpUri-10="https://zenodo.org/record/4398039/files/figure.png" httpUri-11="https://zenodo.org/record/4398041/files/figure.png" httpUri-12="https://zenodo.org/record/4398043/files/figure.png" httpUri-2="https://zenodo.org/record/4398023/files/figure.png" httpUri-3="https://zenodo.org/record/4398025/files/figure.png" httpUri-4="https://zenodo.org/record/4398027/files/figure.png" httpUri-5="https://zenodo.org/record/4398029/files/figure.png" httpUri-6="https://zenodo.org/record/4398031/files/figure.png" httpUri-7="https://zenodo.org/record/4398033/files/figure.png" httpUri-8="https://zenodo.org/record/4398035/files/figure.png" httpUri-9="https://zenodo.org/record/4398037/files/figure.png" pageId="1" pageNumber="439">Figs 1–71</figureCitation>
)
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<treatmentCitation id="9F041095160D753056E3FC5EB0BDBDBB" author="Gillies, M. T." box="[151,453,939,963]" page="125" pageId="1" pageNumber="439" year="1990">
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<emphasis id="2CD1EA96160D753056E3FC5EB077BDBB" box="[151,271,939,963]" italics="true" pageId="1" pageNumber="439">Demoulinia</emphasis>
<bibRefCitation id="7A344B75160D75305761FC5EB0BDBDBB" author="Gillies, M. T." box="[277,453,939,963]" pageId="1" pageNumber="439" pagination="97 - 128" refId="ref10174" refString="Gillies, M. T. (1990) A revision of the African species of Centroptilum Eaton (Baetidae, Ephemeroptera). Aquatic Insects, 12 (2), 97 - 128. https: // doi. org / 10.1080 / 01650429009361395" type="journal article" year="1990">Gillies 1990: 125</bibRefCitation>
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(imago, larva).
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<typeStatus id="C11E8826160D753056E3FC1CB1B4BA79" box="[151,204,1001,1025]" pageId="1" pageNumber="439">Type</typeStatus>
species:
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<taxonomicName id="D9A54D07160D75305758FC1DB3E2BA79" authority="Demoulin 1970" authorityName="Demoulin" authorityYear="1970" box="[300,666,1000,1025]" class="Insecta" family="Baetidae" genus="Centroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="439" phylum="Arthropoda" rank="species" species="crassi">
<emphasis id="2CD1EA96160D75305758FC1DB08DBA78" box="[300,501,1000,1024]" italics="true" pageId="1" pageNumber="439">Centroptilum crassi</emphasis>
<bibRefCitation id="7A344B75160D7530578FFC1DB3E2BA79" author="Demoulin, G." box="[507,666,1000,1025]" pageId="1" pageNumber="439" pagination="24 - 170" refId="ref9866" refString="Demoulin, G. (1970) Ephemeroptera des faunes ethiopienne et malgache. In: Hanstrom, B., Brinck, P. &amp; Rudebeck, G. (Eds.), South African Animal Life (Results of the Lunds University Expedition in 1950 - 1951), 14, pp. 24 - 170." type="journal article" year="1970">Demoulin 1970</bibRefCitation>
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(objective synonym of 
<taxonomicName id="D9A54D07160D753055FEFC1DB58ABA79" authority="Crass 1947" authorityName="Crass" authorityYear="1947" box="[906,1266,1000,1025]" class="Insecta" family="Baetidae" genus="Centroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="439" phylum="Arthropoda" rank="species" species="pulchrum">
<emphasis id="2CD1EA96160D753055FEFC1DB500BA78" box="[906,1144,1000,1024]" italics="true" pageId="1" pageNumber="439">Centroptilum pulchrum</emphasis>
<bibRefCitation id="7A344B75160D7530520AFC1CB58ABA79" author="Crass, R. S." box="[1150,1266,1001,1025]" pageId="1" pageNumber="439" pagination="37 - 100" refId="ref9802" refString="Crass, R. S. (1947) The May-flies (Ephemeroptera) of Natal and the Eastern Cape. Annals of the Natal Museum, 11 (1), 37 - 100." type="journal article" year="1947">Crass 1947</bibRefCitation>
</taxonomicName>
, nom. praeocc.).
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