Taxonomy and evolution of asymmetric male genitalia in the subgenus Ashima Chen (Diptera: Drosophilidae: Phortica Schiner), with descriptions of seven new species Author Toda, Masanori J. Hokkaido University Museum, Hokkaido University, Sapporo 060 - 0810, Japan Author Bänziger, Hans Department of Entomology and Plant Pathology, Faculty of Agriculture, Chiang Mai University, Chiang Mai 50200, Thailand hans. banziger @ cmu. ac. th; https: // orcid. org / 0000 - 0001 - 8913 - 3034 Author Sati, Pradeep C. Systematic & Cytogenetic Laboratory, Department of Zoology and Biotechnology, HNB Garhwal University, Srinagar-Garhwal, 246174, Uttarakhand, India. & satipradeep 28 @ gmail. com; https: // orcid. org / 0000 - 0001 - 6809 - 8575 Author Fartyal, Rajendra S. Systematic & Cytogenetic Laboratory, Department of Zoology and Biotechnology, HNB Garhwal University, Srinagar-Garhwal, 246174, Uttarakhand, India. & rs. fartyal @ hnbgu. ac. in; https: // orcid. org / 0000 - 0002 - 8844 - 6263 Author Suwito, Awit Zoology Division (Museum Zoologicum Bogoriense), Research Center for Biology-LIPI, Jl. Raya Jakarta-Bogor Km. 46, Cibinong 16911, West Java, Indonesia. awitswt @ yahoo. com Author Katoh, Toru Department of Biological Sciences, Faculty of Science, Hokkaido University, Sapporo 060 - 0810, Japan t _ katoh @ sci. hokudai. ac. jp; https: // orcid. org / 0000 - 0003 - 3376 - 8545 text Zootaxa 2020 2020-06-08 4789 1 1 54 journal article 21673 10.11646/zootaxa.4789.1.1 c12aaed8-264c-459c-b1aa-09eaeb686670 1175-5326 3884610 311A27A2-A053-4D20-A43E-EF57DBE55166 18) Phortica ( Ashima ) watabei Sati & Fartyal , sp. nov. ( Fig. 5 ) Diagnosis. Apical expansion of arista 1/4 as wide as long, apically tapering and pointed ( Fig. 5A ); fore and mid tibiae with dark gray rings proximally and distally (ch.17,18-1); posterior postgonite slightly curved outward, 1/2 as long as anterior one on the same lateral lobe; 2 anterior postgonites on the other lateral lobe widely divergent ( Fig. 5E ). Description (not repeating characters common to P . brachychaeta ). Supracervical setae 7–10. Cibarial, medial sensilla approximately 7 per side; posterior sensilla approximately 7 per side. All tarsi with gray tarsomeres IV and V; all tarsomere Is slightly longer than 3 succeeding tarsomeres together. Measurements ( holotype / 1♂ paratype , in mm): BL = (n/a)/2.06, ThL = 1.27/1.42, WL = 2.50/2.52, WW = 1.13/1.14. Indices ( holotype / 1♂ paratype , in ratio): FW/HW = 0.41/0.48, ch/o = 0.11/0.09, prorb = 1.17/1.43, rcorb = 0.83/0.53, vb = (n/a)/0.36, dcl = 0.57/0.42, presctl = (n/a)/0.51, sctl = 1.20/1.00, sterno = 0.88/0.92, orbito = 2.00/2.37, dcp = 0.18/0.27, sctlp = 1.50/1.08, C = 2.60/2.31, 4c = (n/a)/1.68, 4v = 2.35/3.11, 5x = 0.75/1.09, ac = 3.10/2.58, M = (n/a)/0.76, C3F = 0.60/0.51. The antisymmetry is observed in the postgonites: in A-type, the left lateral lobe bears 2 anterior postgonites, and the right lobe 1 posterior and 1 anterior postgonites; but in B-type, vice versa ( Fig. 5E ). FIGURE 5. Phortica ( Ashima ) watabei Sati & Fartyal , sp. nov. (♂ paratype). A, Head (frontal view); B, abdominal sternites; C, periphallic organs (caudal view); D, E, phallic organs (lateral and ventral views, respectively; B-type). Scale bars: 0.1 mm in C–E; 0.5 mm in A, B. Holotype . ♂ (A-type) (DZHNBGU), “ Simli ( Narainbagar ), Chamoli Distr. , Uttarakhand , India , 5.ix.2010 , P.C. Sati ”. Paratype . India : 1♂ (B-type), same data as the holotype ( SEHU ) . Distribution. India ( Uttarakhand ). Remarks. This species is unique in having the fore and mid tibiae with gray color rings (ch.17,18-1) within the foliiseta subgroup, and somewhat resembles P . brachychaeta in the morphology of the aedeagal sheath and postgonites, but can be distinguished from it by the diagnostic characters. Etymology. Patronym, in honor of Emeritus Prof. Hideaki Watabe, Hokkaido University of Education, who has long contributed to the taxonomy of Asian drosophilids.