Wasp-mimicking soldier flies of the Australian Region: revision of Ampsalis Walker, Elissoma White and Lagenosoma Brauer (Stratiomyidae: Clitellariinae) Author Winterton, Shaun L. text Zootaxa 2023 2023-02-24 5246 1 1 63 http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN journal article 56506 10.11646/zootaxa.5246.1.1 18dd960f-0b84-4056-a916-b7438813f724 1175-5326 7673215 BDE8C45B-4F03-403D-8D57-4EFC1584BFE8 Key to Clitellariinae genera of the Oriental and Australian regions (including the Indo-Malayan transition zone) 1 . Notopleural spine present............................................................................... 2 . - Notopleural spine absent............................................................................... 4 . 2 . Apical stylus of antenna with dense and elongate setae (widely distributed from Australia to China ; a single Afrotropical species)............................................................................. Nigritomyia Bigot, 1877 . - Apical stylus of antenna bare or pubescent, without elongate setae.............................................. 3 . 3 . Abdomen broadly rounded ( Indonesia ; widely distributed throughout Oriental and Palaearctic regions).................................................................................................. Clitellaria Meigen, 1800 . - Abdomen elliptical ( Indonesia )...................................................... Anoamyia Lindner, 1935 . 4 . Head relatively flattened dorsoventrally in profile, longer than high............................................. 5 . - Head broadly rounded, or taller and long in profile........................................................... 6 . 5 . Face flat, not produced beyond eye margin in lateral view; antenna short and aristate; wing extends well beyond end of abdomen ( New Zealand )................................................................. Dysbiota Lindner, 1958 . - Face produced anteriorly around base of antenna with a triangular ventral tooth-like projection; antennae elongate, nearly all flagellomeres of equal size and not aristate; wing not extending far beyond end of abdomen ( Indonesia , Papua New Guinea )........................................................................... Caenocephaloides Strand, 1898 . 6 . Scutellum without spines............................................................................... 7 . - Scutellum with marginal spines present.................................................................... 9 . 7 . Abdomen round; antenna relatively short, similar to head length; flagellum tapered apically; body mostly bright orange ( Indonesia , Papua New Guinea , India ).......................................................... Ruba Walker, 1859 . - Abdomen ovoid to elongate; antenna usually much longer than head, flagellum uniform in width along length; body colour variable............................................................................................. 8 . 8 . Abdomen ovoid to petiolate (more so in male), anterior segments (1–3) much narrower than posterior segments (4–8) (e.g., Fig. 33 ); face frequently protruding and observable in lateral view ( Australia ).................... Lagenosoma Brauer, 1882 . - Abdomen ovoid, never petiolate ( Fig. 34D ); face not protruding and barely observable in lateral view ( China , India , Indonesia ).......................................................................... Eudmeta Wiedemann, 1830 . 9 . Wing with cross-vein r–m absent; small, predominantly black species with wings folded over abdomen ( Australia , Papua New Guinea ).......................................................................... Octarthria Brauer, 1882 . - Wing with cross-vein r–m present; body size variable and wings usually straight.................................. 10 . 10 . Flagellum with flagellomeres 4–7 short and strongly tapered, flagellomere 8 very narrow and elongate with terminal style; individuals with black purplish colouration ( Australia )..................................... Geranopus White, 1916 . - Flagellum elongate with flagellomeres uniformly cylindrical, variable in length and shape, lacking hair-like terminal style..................................................................................................... 11 . 11 . Scutellar spines greatly elongate (longer than scutellum length) and strongly divergent in orientation ( Indonesia , Malaysia , Thailand , India , Laos )......................................................... Campeprosopa Macquart, 1850 . - Scutellar spines shorter than scutellum length, sometimes minute, parallel or only moderately divergent in orientation… 12 ... 12. Wing vein M 4 emerging before discal cell ( China , Indonesia , Malaysia , India , Papua New Guinea ; also found in Neotropical and Nearctic regions).......................................................... Cyphomyia Wiedemann, 1819 . - Wing vein M 4 emerging from discal cell.................................................................. 13 . 13. Eyes pilose......................................................................................... 14 . - Eyes not pilose...................................................................................... 15 . 14 . Body black and yellow with extensive yellow setal pile, excellent Vespidae mimics; antenna much longer than head; all flagellomeres similar in shape and width, flagellomeres 1–7 with circular sensory pits ( Australia )... Syndipnomyia Kertész, 1921 . - Body dark in colouration; antenna similar length to head; basal flagellomeres much thicker than distal flagellomeres, flagellomeres 1–3 with circular sensory pits (widely distributed except Australasia)................. Adoxomyia Kertész, 1907 . 15 . Face below antennae flat or rounded, without ‘nose’-like process directed anteroventrally (e.g., Fig. 25E–N ) ( Australia )....................................................................................... Elissoma White, 1916 . - Face below antennae with distinct ‘nose’-like anteroventrally directed process ( Indonesia , Australia ) (e.g., Fig. 25A–D )........................................................................... Ampsalis Walker, 1859 sensu stricto .