Laboulbeniomycetes (Fungi, Ascomycota) of Denmark
Author
Santamaria, Sergi
Unitat de Botànica, Departament de Biologia Animal, de Biologia Vegetal i d’Ecologia, Facultat de Biociències, Universitat Autònoma de Barcelona, 08193 Cerdanyola del Vallès (Barcelona), Spain.
sergi.santamaria@uab.cat
Author
Pedersen, Jan
Natural History Museum of Denmark (Zoological Museum), University of Copenhagen, Universitetsparken 15, 2100 København Ø, Denmark.
japedersen@snm.ku.dk
text
European Journal of Taxonomy
2021
2021-11-26
781
1
425
http://dx.doi.org/10.5852/ejt.2021.781.1583
journal article
10.5852/ejt.2021.781.1583
2118-9773
5828924
Laboulbenia notiophili
Cépède & F.Picard
MB#148452
Fig. 38E–G
Bulletin Scientifique de la
France
et de la
Belgique
42: 259 (
Cépède & Picard 1908
)
. –
Type
: [Type lost!] “Sur les élytres, le prothorax et la tête de
Notiophilus semipunctatus
F.
,
Mortain
(
Manche
). Exemplaires recueillis par M. l’abbé
Pasquet
”. [
France
]
Laboulbenia baetica
Balazuc, Espadaler & Girbal (
Balazuc
et al.
1982: 416
)
[MB#114237]
Laboulbenia blanchardii
Cépède
(
Cépède 1914: 373
; as
L. blanchardi
) [MB#222483]
Laboulbenia cymindicola
Speg. (
Spegazzini 1914: 37
)
[MB#250379]
Laboulbenia pulchella
Speg. (
Spegazzini 1914: 39
)
[MB#145851]
Diagnostic features
Thallus dark except for cell I and, sometimes, cell V. Cell V obtriangular, small, half or less of the length of cell IV, not connected with cell III. Outer appendage unbranched, with the lower cells (including the basal cell) variably darkened, especially towards the dorsal margin. Inner appendage consisting of two branches directly arising from the basal cell, terminated by solitary or a cluster of slender antheridia (
Fig. 38F
, an). [Detailed descriptions:
Weir 1994
;
Majewski 1994b
;
Santamaria 1998
]
Distribution and hosts
Widespread in Europe and North Africa on Col.
Carabidae
of the genera
Notiophilus
,
Cymindis
,
Demetrias
,
Dromius
,
Pseudotrechus
Rosenhauer, 1856
,
Philorhizus
,
Metadromius
,
Calodromius
Reitter, 1905
,
Ophionea
? Klug, 1821,
Homethes
? Newman, 1842 and
Aeptomus
? Found in
Europe
:
Andorra
,
Austria
,
Bulgaria
,
Finland
,
France
,
Germany
,
Hungary
,
Italy
,
Norway
,
Poland
,
Romania
,
Russia
,
Spain
,
Sweden
,
Switzerland
,
United Kingdom
, former
Yugoslavia
(
Santamaria
et al.
1991
),
Portugal
(
Santamaria 1992b
),
Belgium
(
De Kesel 1998
),
Czech Republic
(
Rossi & Máca 2006
),
Denmark
(
Høier 2009
),
the Netherlands
(
Haelewaters
et al.
2012
), and
Africa
:
Algeria
and
Morocco
(
Santamaria
et al.
1991
).
Collections examined from
Denmark
On
Demetrias imperialis
(Germar, 1824)
(Col.
Carabidae
)
DENMARK
–
Lolland, Falster, Møn
(LFM)
•
Vest
for
Bandholm
;
54°50.303′ N
,
11°28.050′ E
; PF57;
3 Jan. 2019
;
JP
1310
;
JP
det.;
ZMUC C-F-123845
. –
Sydsjaelland
(SZ)
•
Østerskov
ved
Langebaek
;
54°59.572′ N
,
12°6.715′ E
; UA19;
3 Jan. 2019
;
JP
1280
;
JP
det.;
ZMUC C-F-123811
.
On
Notiophilus aestuans
Dejean, 1826
(Col.
Carabidae
)
DENMARK
–
Nordøstsjaelland
(NEZ)
•
Vest
for
Trørød
;
55°50.223′ N
,
12°31.185′ E
; UB49;
13 Nov. 2016
;
JP
141
;
JP
det.;
ZMUC C-F-122618
.
On
Notiophilus biguttatus
(Fabricius, 1779)
(Col.
Carabidae
)
DENMARK
–
Lolland, Falster, Møn
(LFM)
•
Krenkerup Haveskov
;
54°46.408′ N
,
11°39.955′ E
; PF77;
12 May 1993
;
S. Langemark
&
O.E. Meyer
900
;
JP
det.;
ZMUC C-F-123398
•
ibid.;
7 Jun. 1993
;
S. Langemark
&
O.E. Meyer
899
;
JP
det.;
ZMUC C-F-123397
•
ibid.;
16 Jun. 1993
;
S. Langemark
&
O.E. Meyer
915
;
JP
det.;
ZMUC C-F-123413
•
ibid.;
8 Jul. 1993
;
S. Langemark
&
O.E. Meyer
905
;
JP
det.;
ZMUC C-F-123403
•
Naesby Strand
;
54°44.625′ N
,
11°4.539′ E
; PF36;
23 Feb. 2014
;
JP
548
;
JP
det.;
ZMUC C-F-123034
•
Vest
for
Bandholm
;
54°50.303′ N
,
11°28.050′ E
; PF57;
3 Jan. 2019
;
JP
1307
;
JP
det.;
ZMUC C-F-123842
. –
Nordøstjylland
(NEJ)
•
Bønderskoven
i
Tofte Skov
;
56°49.955′ N
,
10°14.981′ E
; NH79;
8–22 Aug. 2013
;
I. Aggerholm
928
;
JP
det.;
ZMUC C-F-123428
•
Engskoven
i
Tofte Skov
;
56°51.525′ N
,
10°15.065′ E
; NJ70;
8–22 Aug. 2013
;
I. Aggerholm
938
;
JP
det.;
ZMUC C-F-123440
•
Havgranerne
i
Tofte Skov
;
56°50.576′ N
,
10°15.552′ E
; NJ70;
19 Sep.–2 Oct. 2013
;
I. Aggerholm
924
;
JP
det.;
ZMUC C-F-123422
. –
Nordøstsjaelland
(NEZ)
•
Brønshøj
;
55°42.234′ N
,
12°30.135′ E
; UB47;
8 May 2016
;
JP
64
;
JP
det.;
ZMUC C-F-122541
•
Gribskov
,
Nederste Koppel
;
56°2.749′ N
,
12°20.077′ E
; UC31;
1–31 May 2014
;
J. Heilmann-Clausen et al.
162
;
P. Jørum
det.;
ZMUC C-F-122641
•
Tokkekøb Hegn
;
55°52.334′ N
,
12°22.229′ E
; UB39;
6 May 2013
;
JP
330
;
JP
det.;
ZMUC C-F-122808
. –
Sydsjaelland
(SZ)
•
Krobaek
i
Sjolte Skov
;
55°10.772′ N
,
12°0.863′ E
; UB11;
17 Feb. 2019
;
JP
1373
;
JP
det.;
ZMUC C-F-123914
•
Østerskov
ved
Langebaek
;
54°59.572′ N
,
12°6.715′ E
; UA19;
3 Jan. 2019
;
JP
1274
;
JP
det.;
ZMUC C-F-123804
.
Fig. 38.
Laboulbenia
spp.
A
–D
.
L. murmanica
Huldén.
A
–C
. Mature thalli showing constriction at septum II–VI/III (arrows).
D
. Immature thallus showing undamaged appendage. –
E–G
.
L. notiophili
Cépède & F.Picard.
E
,
G
. Mature
thalli.
F
. Inner appendage in detail showing antheridia (an) and basal cell of inner appendage (bc ia). –
H–I
.
L. olisthopi
Speg.
H
. Mature
thallus.
I
. Two immature thalli with labelled cell V. Scale bars: A–E, G–I = 50 µm; F = 25 µm. Photographs from slides ZMUC C-F-123145 (A–B), ZMUC C-F-123271 (C), ZMUC C-F-123651 (D), ZMUC C-F-122541 (E–F), ZMUC C-F-122545 (G), ZMUC C-F-122637 (H–I).
On
Notiophilus germinyi
Fauvel, 1863
(Col.
Carabidae
)
DENMARK
–
Nordøstjylland
(NEJ)
•
Bønderskoven
i
Tofte Skov
;
56°49.955′ N
,
10°14.981′ E
; NH79;
8–22 Aug. 2013
;
I. Aggerholm
929
;
JP
det.;
ZMUC C-F-123429
.
On
Notiophilus palustris
(Duftschmid, 1812)
(Col.
Carabidae
)
DENMARK
–
Lolland, Falster, Møn
(LFM)
•
Vest
for
Bandholm
;
54°50.303′ N
,
11°28.050′ E
; PF57;
3 Jan. 2019
;
JP
1308
;
JP
det.;
ZMUC C-F-123843
. –
Nordøstjylland
(NEJ)
•
Bønderskoven
i
Tofte Skov
;
56°49.955′ N
,
10°14.981′ E
; NH79;
8–22 Aug. 2013
;
I. Aggerholm
942
;
JP
det.;
ZMUC C-F-123446
. –
Nordøstsjaelland
(NEZ)
•
Asserbo Plantage
;
56°1.984′ N
,
12°0.817′ E
; UC11;
28 Apr. 2019
;
JP
1456
;
JP
det.;
ZMUC C-F-124007
. –
Sydsjaelland
(SZ)
•
Holmegårds Mose
;
55°17.685′ N
,
11°47.928′ E
; PG73;
31 Mar. 2019
;
JP
1416
;
JP
det.;
ZMUC C-F-123962
•
Østerskov
ved
Langebaek
;
54°59.572′ N
,
12°6.715′ E
; UA19;
3 Jan. 2019
;
JP
1275
;
JP
det.;
ZMUC C-F-123805
.
On
Notiophilus rufipes
Curtis, 1829
(Col.
Carabidae
)
DENMARK
–
Nordøstsjaelland
(NEZ)
•
Nordøst
for
Bispebjerg Station
;
55°42.508′ N
,
12°32.786′ E
; UB47;
18 Aug. 2016
;
JP
114
;
JP
det.;
ZMUC C-F-122591
•
Rådvad
;
55°48.338′ N
,
12°33.589′ E
; UB48;
13 Nov. 2016
;
JP
126
;
JP
det.;
ZMUC C-F-122603
•
Vanløse
;
55°41.443′ N
,
12°28.058′ E
; UB47;
15 Jan. 2018
;
J. Søgaard Hansen
995
;
J. Søgaard Hansen
det.;
ZMUC C-F-123502
.
On
Paradromius linearis
(Olivier, 1795)
(Col.
Carabidae
)
DENMARK
–
Fyn
(F)
•
Bagenkop
;
54°45.201′ N
,
10°40.616′ E
; PF06;
5 Oct. 2016
;
JP
149
;
JP
det.;
ZMUC C-F-122628
•
Gyldenstens
inddaemmede
Strand
;
55°34.447′ N
,
10°7.211′ E
; NG75;
2 Apr. 2014
;
JP
694
;
JP
det.;
ZMUC C-F-123183
. –
Lolland, Falster, Møn
(LFM)
•
Møns Klint
,
Syd
;
54°56.985′ N
,
12°32.270′ E
; UA49;
3 Aug. 2019
;
JP
1487
;
JP
det.;
ZMUC C-F-124072
•
Ulvshale
;
55°2.636′ N
,
12°15.638′ E
; UB20;
26 May 2018
;
JP
1110
;
JP
det.;
ZMUC C-F-123628
. –
Nordøstsjaelland
(NEZ)
•
Eriksholm Skov
;
55°41.771′ N
,
11°47.912′ E
; PG77;
4 Nov. 2017
;
JP
737;
JP
det.;
ZMUC C-F-123230
•
Nord
for
Frederikssund
;
55°51.404′ N
,
12°3.534′ E
; UB17;
14 Dec. 2013
;
R. Aagaard Jensen
406
;
JP
det.;
ZMUC C-F-122890
•
Syd
for
Veddelev
;
55°40.346′ N
,
12°4.915′ E
; UB17;
14 Dec. 2013
;
R. Aagaard Jensen
335
;
JP
det.;
ZMUC C-F-122814
•
Vest
for
Trørød
;
55°50.223′ N
,
12°31.185′ E
; UB49;
13 Nov. 2016
;
JP
138
;
JP
det.;
ZMUC C-F-122614
. –
Nordvestjylland
(NWJ)
•
Nord
for
Landting
ved
Vinderup
;
56°30.049′ N
,
8°45.384′ E
; MH86;
19 Jan. 2015
;
JP
572
;
JP
det.;
ZMUC C-F-123058
. –
Nordvestsjaelland
(NWZ)
•
Bognaes Skov
på
Tuse Naes
;
55°44.966′ N
,
11°45.817′ E
; PG78;
10 Dec. 2013
;
JP
878
;
JP
det.;
ZMUC C-F-123375
•
Halleby Å ved
Bjerge Sydstrand
;
55°33.194′ N
,
11°10.079′ E
; PG35;
24 Oct. 2018
;
JP
1209
;
JP
det.;
ZMUC C-F-123735
. –
Sydsjaelland
(SZ)
•
Østerskov
ved
Langebaek
;
54°59.572′ N
,
12°6.715′ E
; UA19;
3 Jan. 2019
;
JP
1279
;
JP
det.;
ZMUC C-F-123810
•
ibid.;
54°59.421′ N
,
12°6.435′ E
; UA19;
19 Aug. 2019
;
JP
1500
;
JP
det.;
ZMUC C-F-124087
.
On
Paradromius longiceps
(Dejean, 1826)
(Col.
Carabidae
)
DENMARK
–
Sydsjaelland
(SZ)
•
Tryggevaelde Å ved Varpelev
;
55°21.055′ N
,
12°15.877′ E
; UB23;
6 Jan. 2018
;
JP
1585
;
JP
det.;
ZMUC C-F-124346
.
On
Philorhizus melanocephalus
(Dejean, 1825)
(Col.
Carabidae
)
DENMARK
–
Østjylland
(EJ)
•
Kjellerup
;
56°17.644′ N
,
9°25.850′ E
; NH23;
16 Feb. 2018
;
JP
976
;
JP
det.;
ZMUC C-F-123483
. –
Fyn
(F)
•
Gyldenstens
inddaemmede
Strand
;
55°34.447′ N
,
10°7.211′ E
; NG75;
2 Apr. 2014
;
JP
695
;
JP
det.;
ZMUC C-F-123184
. –
Lolland, Falster, Møn
(LFM)
•
Vest
for
Bandholm
;
54°50.303′ N
,
11°28.050′ E
; PF57;
3 Jan. 2019
;
JP
1311
;
JP
det.;
ZMUC C-F-123846
. –
Nordøstsjaelland
(NEZ)
•
Ejby Ådal
;
55°42.292′ N
,
11°50.465′ E
; PG77;
16 Feb. 2019
;
L. Kjaer-Thomsen
&
L. Thomas
1395
;
JP
det.;
ZMUC C-F-123938
•
Faergelunden
;
55°50.540′ N
,
12°1.735′ E
; UB19;
15 Jun. 2013
;
JP
229
;
JP
det.;
ZMUC C-F-122710
. –
Nordvestsjaelland
(NWZ)
•
Sonnerup Skov
;
55°56.643′ N
,
11°34.213′ E
;
PH60;
17 May 2019
;
JP
1468
;
JP
det.;
ZMUC C-F-124048
•
Stold Skydeterraen
;
55°45.124′ N
,
11°17.634′ E
; PG48;
25 May 2014
;
JP
377
;
JP
det.;
ZMUC C-F-122860
•
Vesterlyng
;
55°44.195′ N
,
11°17.276′ E
; PG47;
9 Feb. 2014
;
JP
643
;
JP
det.;
ZMUC C-F-123131
. –
Sydsjaelland
(SZ)
•
Tryggevaelde Å ved Varpelev
;
55°21.055′ N
,
12°15.877′ E
; UB23;
6 Jan. 2018
;
JP
965
;
JP
det.;
ZMUC C-F-123472
. –
Vestjylland
(WJ)
•
Velling
;
56°3.078′ N
,
8°18.596′ E
; MH51;
16 Dec. 2018
;
JP
1258
;
JP
det.;
ZMUC C-F-123786
.
On
Philorhizus sigma
(Rossi, 1790)
(Col.
Carabidae
)
DENMARK
–
Bornholm
(B)
•
Dueodde
;
54°59.442′ N
,
15 4.510′ E
; WA09;
2 Jul. 2007
;
JP
69
;
JP
det.;
ZMUC C-F-122545
. –
Østjylland
(EJ)
•
Lindholm Hoved
i
Mossø
;
56°2.772′ N
,
9°42.809′ E
; NH41;
18 Jan. 2015
;
JP
&
H. Liljehult
516
;
JP
det.;
ZMUC C-F-122999
•
Vest
for
Ørnsø
ved
Silkeborg
;
56°9.126′ N
,
9°30.588′ E
; NH32;
24 Nov. 2019
;
JP
1546
;
JP
det.;
ZMUC C-F-124307
. –
Fyn
(F)
•
Gyldenstens
inddaemmede
Strand
;
55°34.447′ N
,
10°7.211′ E
; NG75;
2 Apr. 2014
;
JP
&
H. Liljehult
696
;
JP
det.;
ZMUC C-F-123185
. –
Lolland, Falster, Møn
(LFM)
•
Vest
for
Bandholm
;
54°50.303′ N
,
11°28.050′ E
; PF57;
3 Jan. 2019
;
JP
1312
;
JP
det.;
ZMUC C-F-123847
. –
Nordøstsjaelland
(NEZ)
•
Strødam
;
55°58.056′ N
,
12°16.273′ E
; UC20;
4 Oct. 2014
;
JP
40
;
JP
det.;
ZMUC C-F-122515
. –
Nordvestjylland
(NWJ)
•
Syd
for
Stårup
;
56°34.792′ N
,
9°5.958′ E
; NH07;
6 Feb. 2015
;
JP
219
;
JP
det.;
ZMUC C-F-122700
. –
Sydjylland
(SJ)
•
Sølsted Mose
;
55°1.829′ N
,
8°50.351′ E
; MF89;
2 Jun. 2013
;
JP
486
;
JP
det.;
ZMUC C-F-122970
. –
Sydsjaelland
(SZ)
•
Bimose
i
Broby Overdrev
;
55°23.921′ N
,
11°35.499′ E
; PG64;
5 Aug. 2014
;
JP
41
;
JP
det.;
ZMUC C-F-122516
•
Denderup Sø
i
Denderup Vaenge
;
55°15.075′ N
,
11°57.366′ E
; PG82;
9 Apr. 2019
;
H. Liljehult
1470
;
JP
det.;
ZMUC C-F-124050
•
Tryggevaelde Å ved Varpelev
;
55°21.055′ N
,
12°15.877′ E
; UB23;
6 Jan. 2018
;
JP
966
;
JP
det.;
ZMUC C-F-123473
.
Remarks
Species of
Laboulbenia
on European
Lebiini
were a true taxonomical problem until
Rossi & Santamaria (2006)
proposed that
L. casnoniae
should be regarded as an exclusively American species occurring only on
Colliuris
De Geer, 1774
(Col.
Carabidae
Lebiinae
). Moreover, all previous European and North African records of
L. casnoniae
should be named with the prioritary name
L. notiophili
, including other species as synonyms (see above).
De Kesel
et al.
(2020)
suggested the need for a molecular analysis to confirm this status, especially for the material on
Demetrias
that could belong to
L. blanchardii
as a separate species.