Systematic revision of the genus Everettia Godwin-Austen, 1891 (Mollusca: Gastropoda: Dyakiidae) in Sabah, northern Borneo Author Liew, Thor-Seng Author Schilthuizen, Menno Author Vermeulen, Jaap Jan text Zoological Journal of the Linnean Society 2009 2009-10-26 157 3 515 550 https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2009.00526.x journal article 10.1111/j.1096-3642.2009.00526.x 0024-4082 5443023 EVERETTIA CORRUGATA CORRUGATA LAIDLAW, 1937 Everettia corrugata Laidlaw, 1937: 179–180 , plate 11, figures 1, 2. Everettia corrugata , Schileyko, 2003: 1360 , figure 1775b, c. Type material: BMNH 1936.6 .16.1, holotype (seen) . Material examined: SABAH – Mount Kinabalu: BOR/ MOL 941, southern slope, 3200 m alt., xi.2001 ; BOR/ MOL 2679, southern slope, 3080 m alt., ix.2003 ; BOR/MOL 2688, southern slope, 2865 m alt., ix.2003 ; BOR/MOL 2866, southern slope, 3330 m alt., x.2003 ; BOR/MOL 2869, southern slope, 3080 m alt., ix.2003 ; BOR/MOL 2874, southern slope, 2865 m alt., ix.2003 ; BOR/MOL 2996, southern slope, 2700 m alt., x.2003 ; BOR/MOL 5275, southern slope, 3270 m alt., i.2005 ; BOR/MOL 5276, southern slope, 2800 m alt., ix.2005 ; BOR/MOL 5277, southern slope, 2944 m alt., xi.2005 ; BOR/MOL 5278, southern slope, 2896 m alt., xii.2004 ; BOR/MOL 5279, southern slope, 3088 m alt., ix.2005 ; BOR/MOL 5281, southern slope, 2911 m alt., xii.2004 ; BOR/MOL 5282, southern slope, 3098 m alt., xii.2004 ; BOR/MOL 5283, southern slope, 3290 m alt., i.2005 ; BOR/ MOL 5284, southern slope, 3211 m alt., i.2005 ; BOR/ MOL 5285, southern slope, 2800 m alt., ix.2005 ; BOR/MOL 5286, southern slope, 3024 m alt., ix.2005 ; BOR/MOL 5287, southern slope, 3088 m alt., ix.2005 ; BOR/MOL 5288, southern slope, 2775 m alt., xii.2004 ; BOR/MOL 5289, southern slope, 3154 m alt., i.2005 ; BOR/MOL 5290, southern slope, 3304 m alt., i.2005 ; BOR/MOL 5293, southern slope, 2896 m alt., ix.2005 ; BOR/MOL 5294, southern slope, 3221 m alt., i.2005 ; BOR/MOL 5291, southern slope, 2578 m alt., xi.2005 ; BOR/MOL 5292, southern slope, 3008 m alt., xi.2005 ; BOR/MOL 5280, southern slope, 2640 m alt., xi.2005 . BOR/MOL 2672, southern slope, 3333 m alt., x.2003 ; JJ 4085, southern slope, Paka, 3000 m alt., iv.2004 ; JJ 5953; SP 12107, southern slope, 2600 m alt., x.2000 ; SP 12108, southern slope, 2600 m alt., x.2000 ; SP 12109, southern slope, 2600 m alt., x.2000 ; SP 12323, southern slope, Liwaagu trail, i.2004 ; SP 12338, southern slope, Paka, 3000 m alt., iv.2004 ; SP 12511, southern slope, 2600 m alt., vii.2005 ; SP 12568, southern slope, Silau-silau, vii.2005 ; SP 12659, southern slope, 2600 m alt., xi.2005 ; SP 12666, southern slope, 2600 m alt., xi.2005 ; SP 12670, southern slope, 2600 m alt., xi.2005 ; SP 12837, southern slope, 2600 m alt., xi.2005 ; SP 12947, southern slope, Maraiparai, v.2007 . BMNH 20080211, southern slope, 3000 m alt.; BMNH 20080212, southern slope, 2600 m alt., viii.1967 ; BMNH 20080213, southern slope, 2600 m alt., viii.1967 ; ZMA 176868, southern slope, 3000 m alt., viii.1937 . Figure 10. Shell surface sculptures (scanning electron micrographs). A, upper surface of Everettia jasilini sp. nov. ; B, underside of E. jasilini sp. nov. ; C, upper surface of Everettia safriei sp. nov. ; D; underside of Everettia paulbasintali sp. nov. ; E, upper surface of E. paulbasintali sp. nov. ; F, underside of Everettia klemmantanica ; G, upper surface on the fourth whorl of E. klemmantanica ; H, upper surface on the second whorl of E. klemmantanica . Scale bar = 100 Mm. Figure 11. Shell surface sculptures (scanning electron micrographs). A, upper surface of Everettia corrugata williamsi ssp. nov. ; B, underside of E. corrugata williamsi ssp. nov. ; C, underside of Everettia lapidini ; D, upper surface of Everettia occidentalis sp. nov. ; E, corrugation at the suture of Everettia corrugata corrugata ; F, upper surface of Everettia jucundior sp. nov. Scale bar = 500 Mm. Diagnostic characteristics: Very strong corrugation near the suture. Description: Shell ( Figs 2A , 11E ): medium-sized, rather thin, greenish brown, moderately to distinctly elevated, outer whorls slightly shouldered below the suture. Periphery rounded. Above the periphery, shell shiny with very weak and irregular wrinkling that grows stronger towards suture, and forms corrugation just below the suture. Below the periphery, shell without or with scattered, inconspicuous, fine spiral grooves. Height up to 11.9 mm ; width up to 19.6 mm ; diameter of the first three whorls 1.3–1.4, 1.1–1.2, and 1.85–2.3 mm , respectively; number of whorls up to five; height aperture up to 7.4 mm ; width aperture up to 10.7 mm . Genitalia ( Fig. 7I ): maximum length from genital opening to the end of dart-sac (before the visible gland tubules) up to 11.5 mm . The penis, dartsac, and vagina with thick muscular walls. P, V arranged near the GO then followed by BC and DS where there is little space in the atrium between the openings of P + V and BC + DS. Long BC about twothirds of total DS length. Animal ( Fig. 8G ): animal head uniform black or grey and this colour extends backward to the end of mantle and downward to the foot-fringe. Some individuals with an ill-defined grey band between tentacles, extended to the end of the tail. Grey smear covering large parts of the animal and extending until the end of the tail. The mantle is covered in sparsely distributed brighter spots. These brighter spots become denser and are agglomerated with the black spots at the last half whorl. Distribution and habitat: Upper montane and coniferous primary forest, 2500–3500 m alt. Sabah : Mount Kinabalu only ( Fig. 9B ). Remarks: This species is endemic to Mount Kinabalu.