Systematic revision of the genus Everettia Godwin-Austen, 1891 (Mollusca: Gastropoda: Dyakiidae) in Sabah, northern Borneo Author Liew, Thor-Seng Author Schilthuizen, Menno Author Vermeulen, Jaap Jan text Zoological Journal of the Linnean Society 2009 2009-10-26 157 3 515 550 https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2009.00526.x journal article 10.1111/j.1096-3642.2009.00526.x 0024-4082 5443023 EVERETTIA JASILINI SP. NOV. Types: MALAYSIA : State of Sabah : Kota Belud District . North-western part of Kinabalu Park. At 2900 m alt. along the trail from Sayap Substation to Nonohan peak ( 6°7 ′ N , 116°43 ′ E ). Collected by T.- S. Liew , J. Lapidin , Dominik , and Jasilin. Date : 25.xi.2005 , holotype , BOR/MOL 5318, 12A; four paratypes , two in BMNH 20080630 and two in SP 13062 . Etymology: This species is named after Jasilin, an experienced mountain guide from Kampung Mesilau, who, during the intensive malacological survey on Mount Kinabalu in 2005, made possible many fieldtrips to remote areas on Mount Kinabalu. Material examined: SABAH – Mount Kinabalu: BOR/ MOL 5311, eastern slope, 3416 m alt., vi.2005 ; BOR/ MOL 5314, eastern slope, 3376 m alt., vi.2005 ; BOR/MOL 5315, eastern slope, 3320 m alt., vi.2005 ; BOR/MOL 5317, eastern slope, 3084 m alt., vi.2005 ; BOR/MOL 5312, western slope, 3008 m alt., xi.2005 ; BOR/MOL 5313, western slope, 2816 m alt., xi.2005 ; BOR/MOL 5316, western slope, 2672 m alt., xi.2005 ; BOR/MOL 5319, western slope, 2816 m alt., xi.2005 . Figure 12. Shells. A, Everettia jasilini sp. nov. ; B, Everettia monticola sp. nov. ; C, Everettia dominiki sp. nov. ; D, Everettia safriei sp. nov. ; E, Everettia planispira sp. nov. Scale bar = 10 mm. Figure 13. Genitalia. A, Everettia jasilini sp. nov. ; B, Everettia dominiki sp. nov. ; C, Everettia safriei sp. nov. Diagnostic characteristics: Fine and regularly placed nodules on the upper and lower surface of the shell. Description: Shell ( Figs 10A, B , 12A ): small, rather thin, yellowish brown, moderately to distinctly elevated, outer whorls slightly shouldered below the suture. Periphery rounded. Above the periphery, shell silky with distinct oblique wrinkling; fine (40¥ magn.) but very distinct nodules are arranged spirally (20–35 rows per whorl) on top of the wrinkling. Below the periphery, shell without or with indistinct, widely spaced spiral rows of nodules (2–4 per mm). Height up to 6.8 mm ; width up to 10.7 mm ; diameter of the first three whorls 0.75–0.8, 0.6–0.65, and 1.0– 1.1 mm , respectively; number of whorls up to four and seven-eighths; height aperture up to 4.3 mm ; width aperture up to 5.5 mm . Genitalia ( Fig. 13A ): maximum length from genital opening to the end of dart-sac (before the visible gland tubules) up to 6 mm . The penis and dart-sac have a thin muscular wall and the vagina has a very thin transparent wall. P, V, GO, BC, and DS arranged around the short genital atrium, with limited space between the openings of each of the genital structures. BC short, about half of total DS length. Animal: the whole animal head is uniformly black. The mantle is black and covered by sparsely distributed tiny bright spots. Distribution and habitat: Upper montane primary and coniferous forest, 3100–3500 m alt. Sabah : Mount Kinabalu, endemic (eastern and northern ridges only; Fig. 9B ). Remarks: Phylogenetic data show that this species is closely related to Everettia safriei .