Diversity and taxonomic review of Leptusa Kraatz (Coleoptera: Staphylinidae: Aleocharinae) from Great Smoky Mountains National Park, U. S. A., with descriptions of four new species Author Park, Jong-Seok Author Carlton, Christopher E. Author Ferro, Michael L. text Zootaxa 2010 2662 1 27 journal article 10.5281/zenodo.199052 a07f4b3c-e602-4b4e-95dc-87fef597d912 1175-5326 199052 Leptusa ( Adoxopisalia ) stocksi Park & Carlton , new species ( Figs. 1 d. 5a – i, Map 1) Holotype . U.S.A. : Tennessee: Sevier co.: ɗ, “TENNESSEE: Sevier Co., GSMNP , Clingmans Dome ATBI Plot. Malaise trap MT 16200110816. 31 Jul–16 Aug 2001. I.C.Stocks”, “ HOLOTYPE , Leptusa stocksi Park and Carlton , des. 2010”. Deposited in the Field Museum, Chicago. Paratypes (n=2). U.S.A. , Tennessee: 2ɗɗ with same data as holotype GSMNP and LSAM (slide mounted specimen missing genitalia). Diagnosis. Leptusa ( Adoxopisalia ) stocksi has similar appearance with L . ( A .) opaca , but it can be distinguished by shorter antennomeres 4 – 7 ( Fig. 5 a), shape of median lobe ( Fig. 5 i), male abdominal tergites VII – VIII without single median elongate tubercle ( Fig. 5 f), and short δ-seta (as long as α-, β-seta) of labial palpi ( Fig. 5 d). Description of male. Length 3.0 – 3.3 mm . Body brown; antennomeres 1 – 3, 11, mouthparts, legs, and abdominal tergites II – V yellowish brown ( Fig. 1 c). Head opaque, pubescent, with microsculpture. Antennomeres 1 – 3 elongate, 4 longer than wide, 5 – 6 subquadrate, 7 – 10 transverse ( Fig. 5 a). Carina on each side of ventral surface of head complete, attaining gular suture. Mouthparts . Labrum transverse, bearing 3 pairs of short setae, and 7 pairs of long setae, a-seta, b-seta, and many pores ( Fig. 5 b). Labium with two distal setae in a longitudinal row, and several pores in median area; a pair of setal pores, 2 pairs of real pores and several pseudopores present in lateral area ( Fig. 5 c). Labial palpi bearing 12 setae (a – h, α – δ) ( Fig. 5 d). Mentum trapezoidal, bearing 4 pairs of main setae (b, u, v, w), a pair of additional setae, and many pores ( Fig. 5 e). Thorax. Pronotum wider than head, and approximately 1.44 times wider than long; pubescent, opaque, with microsculpture. Mesoventrite with distinct reticulate microsculpture. Metaventrite with setigerous punctures. Elytra opaque, wider than pronotum and abdomen, approximately 1.58 times longer than wide, approximately 1.56 times longer than pronotum, and latero-posterior margin emarginate. Hind wings present. Abdomen . Abdomen somewhat glossy. Abdominal tergite VI approximately 1.99 wider than long. Male abdominal tergite VIII with posterior margin emarginate, bearing 4 pairs of main setae, and many additional short setae and pores ( Fig. 5 f). Male abdominal sternite VIII with posterior margin produced, bearing 6 pairs of main setae, and many additional short setae and pores ( Fig. 5 g). Genitalia . Paramere as in Fig. 5 h. Median lobe as in Fig. 5 i. Female. Unknown. Distribution. USA : Tennessee. Locations where L. ( A. ) stocksi has been collected in GSMNP as in Map 1. Etymology. This species is named in honor of Ian Stocks (Clemson University, Clemson, SC), collector of the type series. Habitat. This species is only known from Clingmans Dome, the highest point in GSMNP. All specimens were collected using a malaise trap during August.