Systematic Revision Of The Phorusrhacidae (Aves: Ralliformes) Author Alvarenga, Herculano M. F. Museu de História Natural de Taubaté. Rua Colômbia, 99, CEP 12030 - 520, Taubaté, SP, Brasil. halvarenga@uol.com.br Author Höfling, Elizabeth Departamento de Zoologia, Instituto de Biociências, Universidade de São Paulo, Caixa Postal 11.294, CEP 05422 - 970, São Paulo, SP, Brazil. text Papéis Avulsos de Zoologia 2003 2003-12-31 43 4 55 91 journal article 10.1590/S0031-10492003000400001 1807-0205 4900701 Genus Titanis Brodkorb, 1963 Type Species – Titanis walleri Brodkorb, 1963 . Included Species – Only the type species. Distribution – Upper Pliocene to Lower Pleistocene of Florida and Texas ( U.S.A. ). Diagnosis – Size similar to that of Phorusrhacos , with a less sturdy tarsometatarsus than that of Devincenzia . The mid-trochlea, in the more distal portion, is spread out onto the sides, differentiating it from the two aforementioned species ( Fig. 22C ), this aspect reminding one of the Mesembriornithinae .