Systematic Revision Of The Phorusrhacidae (Aves: Ralliformes) Author Alvarenga, Herculano M. F. Museu de História Natural de Taubaté. Rua Colômbia, 99, CEP 12030 - 520, Taubaté, SP, Brasil. halvarenga@uol.com.br Author Höfling, Elizabeth Departamento de Zoologia, Instituto de Biociências, Universidade de São Paulo, Caixa Postal 11.294, CEP 05422 - 970, São Paulo, SP, Brazil. text Papéis Avulsos de Zoologia 2003 2003-12-31 43 4 55 91 journal article 10.1590/S0031-10492003000400001 1807-0205 4900701 Genus Devincenzia Kraglievich, 1932 Devincenzia Kraglievich, 1932:323 , 338. Onactornis Cabrera, 1939:15 ; syn. n. TABLE 5. Measurements of Phorusrhacos longissimus (cm). Numbers in brackets are estimates based on incomplete bones.

MLP-	BMNH-	BMNH-	BMNH-
118	A529	A530	A684
Skull*
Total length	–	(65)	–	–
Width at base	–	(30)	–	–
Height of maxilla	–	(25)	–	–
Mandible
Total length	–	55	–	–
Length of symphysis	(16)	17.9	–	–
Height- base of the symphysis	5.3	6.2	5.7	5.4
Width- base of symphysis	7.0	6.7	6.6	6.0
BMNH-	BMNH-	BMNH-	MLP-	MLP-	AMNH-	AMNH-
A581	A531	A545	131	76V.1011	9497	9146
Femur
Total length	–	–	–	–	–	–	(31.0)
Width middle of diaphysis	–	–	–	–	–	–	3.62
Maximum distal width	–	–	–	–	–	–	9.2
Tibiotarsus
Total length	–	–	–	–	–	–	(50.0)
Smallest width of diaphysis	–	–	–	–	–	–	3.9
Maximum distal width	–	–	–	–	–	–	6.2
Tarsometatarsus
Total length	–	–	–	38.5	(40.0)	37.0	–
Proximal width	–	–	–	8.0	(8.4)	–	–
Width- middle of diaphysis	–	–	–	3.7	3.7	3.6	–
Width at distal foramen level	6.0	7.1	6.5	6.4	6.5	6.3	–
Width of the middle trochea	2.9	–	3.5	3.3	3.6	3.4	–

* Measurements from Ameghino (1895) Type Species – Devincenzia gallinali Kraglievich, 1932 , junior synonym of Devincenzia pozzi (Kraglievich, 1931) . Included Species – Only the type species. Distribution – Upper Miocene to the Lower Pliocene of Argentina and Uruguay . Diagnosis – They are the biggest of the known Phorusrhacinae , with a height comparable to that of the largest Brontornithinae , being, however, lighter. The skull is about 65 cm long, equivalent to that of Phorusrhacos , this being possibly flattened dorso-ventrally ( Fig. 32A ). The tarsometatarsus is substantially sturdier than that of Phorusrhacos ( Figs. 2F and 22A ). Remarks – The general appearance of the tarsometatarsus of Devincenzia (MNHN-189, Fig. 2F ), appears to indicate an intermediate position between the Phorusrhacinae and Brontornithinae , as observed by Kraglievich (1932) himself. However, the appearance of the bony cortex makes it clear that one is dealing with a not yet completely adult individual. As happens with several present-day Ralliformes , especially the Rallidae and Cariamidae , besides other groups of birds, it is normal that the proximal metaphysary portion of the tarsometatarsus reaches, in youngsters, a larger width than that in the adults, associated to a shorter length of the bone. Consequently, one supposes that the tarsometatarsus of Devincenzia , in the adult phase, might be a little longer and with the proximal region slightly narrower than appears in the refered specimen. The origin of this tarsometatarsus (MNHN-189), type of Devincenzia gallinali Kraglievich, 1932 , and, consequently, its geological age, are unknown. From the appearance of the bone (and certainly the remains of the encrusted substract), Kraglievich supposed it to have originated from the Arroyo Roman, Rio Negro , Uruguay , and points out the difference of color, especially when compared with the bones coming from Patagonia, without, however, comparing it with specimens from the Huayquerian of north Argentina , studied years before by he himself (1931).