Systematic Revision Of The Phorusrhacidae (Aves: Ralliformes) Author Alvarenga, Herculano M. F. Museu de História Natural de Taubaté. Rua Colômbia, 99, CEP 12030 - 520, Taubaté, SP, Brasil. halvarenga@uol.com.br Author Höfling, Elizabeth Departamento de Zoologia, Instituto de Biociências, Universidade de São Paulo, Caixa Postal 11.294, CEP 05422 - 970, São Paulo, SP, Brazil. text Papéis Avulsos de Zoologia 2003 2003-12-31 43 4 55 91 journal article 10.1590/S0031-10492003000400001 1807-0205 4900701 Genus Paraphysornis Alvarenga, 1993 Type Species – Physornis brasiliensis Alvarenga, 1982 . Included Species – Only the type species. TABLE 3. Measurements of Physornis fortis (cm). Numbers in brackets are estimates on incomplete bones.

Mandible	FM-P13340	FM-P13619
Length of the mandible on the dorsal surface	(11.0)	11.3
Width of the base of the symphysis	7.0	(7.5)
Height at the base of the symphysis	(5.9)	–
Atlas	FM-P13340
Condyloid fossa (maximum width)	3.82
Femur*
Smallest transverse diameter of the diaphysis	5.8
Maximum distal width	14.8
Tarsometatarsus	MACN-A52-185
Maximum proximal width	10.5
Proximal dorsoventral diameter	6.7
Width of the diaphysis on the level of the fracture (Fig. 13)	5.4
Falanx 1, digit IV, left foot	MACN-A52-188
Length of the axis	6.3
Maximum proximal width	4.5
Maximum distal width	(3.2)

* Measurements from Loomis (1914) of the specimen (w/n.) in Amherst College, Massachusetts. Distribution – Upper Oligocene or the Lower Miocene of Southeast Brazil . Diagnosis – Maybe the smallest of the Brontornithinae , the size being comparable to that of a smaller-sized Brontornis burmeisteri ( Figs. 1B and 2B ). The mandibular symphysis is longer and narrower than in the remaining Brontornithinae ( e . g . Alvarenga, 1993: Fig. 1 ), being, however, proportionally wider and shorter than in the Phorusrhacinae . The cotyles of the tarsometatarsus are slightly quadrangular, especially the inner one ( e . g . Alvarenga, 1993: Fig. 2 ).The lateral edge of the hypotarsus expands until very near to the lateral part of the lateral cotyle, not forming the lateral crest characteristic of Physornis .