<document id="5C6D37E0A8D2A86AD487AE0B56DF358B" ID-DOI="10.11646/zootaxa.4021.2.10" ID-GBIF-Dataset="73fffbcd-7c72-45ab-bce7-43db46d82db4" ID-ISSN="1175-5326" ID-Zenodo-Dep="245568" ID-ZooBank="ABAB07DA-FAA4-40D6-AA58-18E160380068" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461276299032" checkinUser="plazi" docAuthor="Volynkin, Anton V., Matov, Alexej Yu., Stüning, Dieter &amp; Behounek, Gottfried" docDate="2015" docId="03CF87C7FFE6FF8A31CCFF5AFBB9469B" docLanguage="en" docName="zt04021p400.pdf" docOrigin="Zootaxa 4021 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Acronicta (Viminia) grumi subsp. bicolor Moore 1881" docType="treatment" docVersion="8" lastPageNumber="400" masterDocId="FFF6FFBFFFE3FF8F315BFFCDFFFE451A" masterDocTitle="On the taxonomy of the genus Acronicta Ochsenheimer, 1816 (Lepidoptera, Noctuidae). I. Redescription of Acronicta grumi (Alphéraky, 1897), with notes on synonymy and its subgeneric placement" masterLastPageNumber="400" masterPageNumber="395" pageNumber="400" updateTime="1698612918568" updateUser="plazi">
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<mods:title id="2184B0F6608EBD7B26D01B1D7DB555E6">On the taxonomy of the genus Acronicta Ochsenheimer, 1816 (Lepidoptera, Noctuidae). I. Redescription of Acronicta grumi (Alphéraky, 1897), with notes on synonymy and its subgeneric placement</mods:title>
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<mods:namePart id="2799A71CD748A7ED8DABB97BA5CA14AA">Volynkin, Anton V.</mods:namePart>
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<mods:namePart id="C8B4302CCCF3F9B2069960044C3B37B0">Matov, Alexej Yu.</mods:namePart>
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<mods:namePart id="C6C4937D0DDC927B3880149496042CB8">Stüning, Dieter</mods:namePart>
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<mods:namePart id="DEEF8AB7E920C6A0396DD32263E3A0FA">Behounek, Gottfried</mods:namePart>
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<mods:date id="47C2147700EBB1FD139B550590F03283">2015</mods:date>
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<treatment id="03CF87C7FFE6FF8A31CCFF5AFBB9469B" ID-DOI="http://doi.org/10.5281/zenodo.6100578" ID-GBIF-Taxon="119639702" ID-Zenodo-Dep="6100578" LSID="urn:lsid:plazi:treatment:03CF87C7FFE6FF8A31CCFF5AFBB9469B" httpUri="http://treatment.plazi.org/id/03CF87C7FFE6FF8A31CCFF5AFBB9469B" lastPageNumber="400" pageId="5" pageNumber="400">
<subSubSection id="C37C655AFFE6FF8A31CCFF5AFE93442D" pageId="5" pageNumber="400" type="nomenclature">
<paragraph id="8BD936D1FFE6FF8A31CCFF5AFC6145A8" blockId="5.[151,927,151,209]" box="[151,927,151,178]" pageId="5" pageNumber="400">
<heading id="D09181BDFFE6FF8A31CCFF5AFC6145A8" bold="true" box="[151,927,151,178]" fontSize="11" level="1" pageId="5" pageNumber="400" reason="1">
<emphasis id="B912EAC3FFE6FF8A31CCFF5AFC6145A8" bold="true" box="[151,927,151,178]" pageId="5" pageNumber="400">
<taxonomicName id="4C664D52FFE6FF8A31CCFF5AFD1545A8" ID-CoL="9NCY" authority="Moore, 1881" authorityName="Moore" authorityYear="1881" box="[151,747,151,178]" class="Insecta" family="Noctuidae" genus="Acronicta" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="400" phylum="Arthropoda" rank="subSpecies" species="grumi" subGenus="Viminia" subSpecies="bicolor">
<emphasis id="B912EAC3FFE6FF8A31CCFF5AFEF345AB" bold="true" box="[151,269,151,177]" italics="true" pageId="5" pageNumber="400">Acronicta</emphasis>
(
<emphasis id="B912EAC3FFE6FF8A3045FF5AFE8345AB" bold="true" box="[286,381,151,177]" italics="true" pageId="5" pageNumber="400">Viminia</emphasis>
) 
<emphasis id="B912EAC3FFE6FF8A30D4FF5AFDCB45AB" bold="true" box="[399,565,151,177]" italics="true" pageId="5" pageNumber="400">grumi bicolor</emphasis>
(Moore, 1881)
</taxonomicName>
syn. &amp; stat. n.
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD936D1FFE6FF8A31CCFF77FE8345CB" blockId="5.[151,927,151,209]" box="[151,381,186,209]" pageId="5" pageNumber="400">
(
<figureCitation id="135D2A54FFE6FF8A31C5FF77FF0045CB" box="[158,254,186,209]" captionStart="FIGURES 1 – 10" captionStartId="2.[151,258,1860,1882]" captionTargetBox="[157,1436,198,1838]" captionTargetId="figure@2.[151,1436,191,1839]" captionTargetPageId="2" captionText="FIGURES 1 – 10. Acronicta spp.: adults. 1, A. grumi, holotype male, Tien Shan (ZISP); 2, A. grumi, male, holotype of A. tiena, [Kyrgyzstan, Kirgizsky Ridge] &quot; Alexandergebirge &quot; (NKMB, photo by W. Mey); 3, A. grumi, female, [Kyrgyzstan, Kirgizsky Ridge] &quot; Alexandergeb. &quot; (ZSM); 4, A. grumi bicolor, syntype of Acronycta bicolor, female, N India, Solun (© NHM) (photo by A. Zilli); 5, A. grumi bicolor, female, N India, Solun (ZISP); 6, A. grumi bicolor, female, China, Tibet (ZSM); 7, A. rumicis, female, West Siberia (AVB); 8, A. rumicis, male, SW Mongolia (AVB); 9, A. indica, male, N India, Sikkim (ZISP); 10, A. megacephala, type species of Subacronicta, European part of Russia, Tula reg. (ZISP)." httpUri="https://zenodo.org/record/245569/files/figure.png" pageId="5" pageNumber="400">Figs 4–6</figureCitation>
, 
<figureCitation id="135D2A54FFE6FF8A3051FF77FED845CB" box="[266,294,186,209]" captionStart="FIGURES 11 – 16" captionStartId="3.[151,258,1951,1973]" captionTargetBox="[187,1431,198,1929]" captionTargetId="figure@3.[151,1436,191,1930]" captionTargetPageId="3" captionText="FIGURES 11 – 16. Acronicta spp.: male genitalia. 11, A. grumi, holotype, NW China, slide Matov 0144; 12, A. grumi, holotype of A. tiena, Kyrgyzstan, slide MB 358 Boursin (photo by W. Mey); 13, A. grumi bicolor, China, slide Stüning 2288 - DS; 14, A. rumicis, East Kazakhstan, slide AV 0931 Volynkin; 15, A. indica, N India, slide Matov 0385; 16, A. digna, Russian Far East, slide AV 1499 Volynkin." httpUri="https://zenodo.org/record/245570/files/figure.png" pageId="5" pageNumber="400">13</figureCitation>
, 
<figureCitation id="135D2A54FFE6FF8A3069FF77FE8845CB" box="[306,374,186,209]" captionStart="FIGURES 17 – 24" captionStartId="4.[151,258,1637,1659]" captionTargetBox="[155,1431,200,1614]" captionTargetId="figure@4.[151,1436,191,1616]" captionTargetPageId="4" captionText="FIGURES 17 – 24. Acronicta spp.: male (17 – 19) and female (20 – 24) genitalia. 17, A. megacephala, type species of Subacronicta, Austria, Vienna, slide AV 1024 Volynkin; 18, A. grumi, aedeagus of holotype, diagnostically important sclerotized elements of vesica; 19, A. grumi, aedeagus of tiena lectotype, diagnostically important sclerotized elements of vesica; 20, A. grumi, Alexandergebirge, slide AV 1459 Volynkin; 21, A. grumi bicolor, China, slide AV 1460 Volynkin; 22, A. grumi bicolor, N India, Solun, slide Matov 0396; 23, A. rumicis, Russia, Altai Mts., slide AV 1501 Volynkin; 24, A. indica, Nepal, slide Stüning 2289 - DS." httpUri="https://zenodo.org/record/245571/files/figure.png" pageId="5" pageNumber="400">21, 22</figureCitation>
)
</paragraph>
<paragraph id="8BD936D1FFE6FF8A31CCFECDFE93442D" blockId="5.[151,1436,256,311]" pageId="5" pageNumber="400">
<taxonomicName id="4C664D52FFE6FF8A31CCFECDFE13440D" authority="Moore, 1881" authorityName="Moore" authorityYear="1881" box="[151,493,256,279]" class="Insecta" family="Noctuidae" genus="Acronycta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B912EAC3FFE6FF8A31CCFECDFEA6440D" box="[151,344,256,279]" italics="true" pageId="5" pageNumber="400">Acronycta bicolor</emphasis>
Moore, 1881
</taxonomicName>
, 
<emphasis id="B912EAC3FFE6FF8A30A1FECDFBF7440D" box="[506,1033,256,279]" italics="true" pageId="5" pageNumber="400">Proceedings of the Zoological Society of London</emphasis>
1881: 332 (
<typeStatus id="54DD8873FFE6FF8A35D7FECDFB3D440D" box="[1164,1219,256,279]" pageId="5" pageNumber="400">Type</typeStatus>
locality: NW 
<collectingCountry id="F3717641FFE6FF8A3405FECDFA69440D" box="[1374,1431,256,279]" name="India" pageId="5" pageNumber="400">India</collectingCountry>
, Punjab, Solun).
</paragraph>
</subSubSection>
<subSubSection id="C37C655AFFE6FF8A31CCFEAFFED847AB" pageId="5" pageNumber="400" type="materials_examined">
<paragraph id="8BD936D1FFE6FF8A31CCFEAFFC1E471E" blockId="5.[151,1436,354,897]" pageId="5" pageNumber="400">
<emphasis id="B912EAC3FFE6FF8A31CCFEAFFE514463" bold="true" box="[151,431,354,377]" pageId="5" pageNumber="400">
<typeStatus id="54DD8873FFE6FF8A31CCFEAFFF314463" box="[151,207,354,377]" pageId="5" pageNumber="400">Type</typeStatus>
material examined
</emphasis>
. Photographs of 
<typeStatus id="54DD8873FFE6FF8A3334FEAFFD2F4463" box="[623,721,354,377]" pageId="5" pageNumber="400" type="syntype">
<emphasis id="B912EAC3FFE6FF8A3334FEAFFD2F4463" bold="true" box="[623,721,354,377]" pageId="5" pageNumber="400">syntypes</emphasis>
</typeStatus>
: 
<specimenCount id="9D60FD58FFE6FF8A33B9FEAFFCBF4463" box="[738,833,354,377]" pageId="5" pageNumber="400" type="female">1 female</specimenCount>
(
<figureCitation id="135D2A54FFE6FF8A3208FEAFFC694463" box="[851,919,354,377]" captionStart="FIGURES 1 – 10" captionStartId="2.[151,258,1860,1882]" captionTargetBox="[157,1436,198,1838]" captionTargetId="figure@2.[151,1436,191,1839]" captionTargetPageId="2" captionText="FIGURES 1 – 10. Acronicta spp.: adults. 1, A. grumi, holotype male, Tien Shan (ZISP); 2, A. grumi, male, holotype of A. tiena, [Kyrgyzstan, Kirgizsky Ridge] &quot; Alexandergebirge &quot; (NKMB, photo by W. Mey); 3, A. grumi, female, [Kyrgyzstan, Kirgizsky Ridge] &quot; Alexandergeb. &quot; (ZSM); 4, A. grumi bicolor, syntype of Acronycta bicolor, female, N India, Solun (© NHM) (photo by A. Zilli); 5, A. grumi bicolor, female, N India, Solun (ZISP); 6, A. grumi bicolor, female, China, Tibet (ZSM); 7, A. rumicis, female, West Siberia (AVB); 8, A. rumicis, male, SW Mongolia (AVB); 9, A. indica, male, N India, Sikkim (ZISP); 10, A. megacephala, type species of Subacronicta, European part of Russia, Tula reg. (ZISP)." httpUri="https://zenodo.org/record/245569/files/figure.png" pageId="5" pageNumber="400">Fig. 4</figureCitation>
), handwritten whitish label &quot; 
<taxonomicName id="4C664D52FFE6FF8A3582FEAFFA624463" box="[1241,1436,354,377]" class="Insecta" family="Noctuidae" genus="Acronycta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B912EAC3FFE6FF8A3582FEAFFA624463" box="[1241,1436,354,377]" italics="true" pageId="5" pageNumber="400">Acronycta bicolor</emphasis>
</taxonomicName>
<typeStatus id="54DD8873FFE6FF8A31CCFE48FF324486" box="[151,204,389,412]" pageId="5" pageNumber="400">Type</typeStatus>
Moore&quot; / printed whitish label &quot;Moore Coll. 94–106&quot; / printed round white label with red ring &quot; 
<typeStatus id="54DD8873FFE6FF8A3459FE48FAC24486" box="[1282,1340,389,412]" pageId="5" pageNumber="400">Type</typeStatus>
&quot; / white printed label with QR-code &quot; 
<collectionCode id="ED77AE14FFE6FF8A308DFE65FDD544A5" box="[470,555,424,447]" pageId="5" pageNumber="400">BMNH</collectionCode>
(E) 1377132&quot; (coll. NHM); 
<specimenCount id="9D60FD58FFE6FF8A323BFE65FC2244A5" box="[864,988,424,447]" pageId="5" pageNumber="400" type="generic">1 specimen</specimenCount>
without body, handwritten whitish label &quot; 
<taxonomicName id="4C664D52FFE6FF8A31F9FE07FE9D44FB" box="[162,355,458,481]" class="Insecta" family="Noctuidae" genus="Acronycta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B912EAC3FFE6FF8A31F9FE07FE9D44FB" box="[162,355,458,481]" italics="true" pageId="5" pageNumber="400">Acronycta bicolor</emphasis>
</taxonomicName>
<typeStatus id="54DD8873FFE6FF8A3031FE07FE6144FB" box="[362,415,458,481]" pageId="5" pageNumber="400">Type</typeStatus>
Moore&quot; / printed whitish label &quot;Moore Coll. 94–106&quot; / printed round white label with red ring &quot; 
<typeStatus id="54DD8873FFE6FF8A31C4FE20FF27471E" box="[159,217,493,516]" pageId="5" pageNumber="400">Type</typeStatus>
&quot; / white printed label with QR-code &quot; 
<collectionCode id="ED77AE14FFE6FF8A3331FE20FD41471E" box="[618,703,493,516]" pageId="5" pageNumber="400">BMNH</collectionCode>
(E) 1377133&quot; (coll. NHM).
</paragraph>
<paragraph id="8BD936D1FFE6FF8A319CFDDDFED847AB" blockId="5.[151,1436,354,897]" pageId="5" pageNumber="400">
<emphasis id="B912EAC3FFE6FF8A319CFDDDFE17473D" bold="true" box="[199,489,528,551]" pageId="5" pageNumber="400">Other material examined</emphasis>
. 
<specimenCount id="9D60FD58FFE6FF8A30A2FDDDFDA4473D" box="[505,602,528,551]" pageId="5" pageNumber="400" type="female">1 female</specimenCount>
, 
<taxonomicName id="4C664D52FFE6FF8A333CFDDDFCD7473D" box="[615,809,528,551]" class="Insecta" family="Noctuidae" genus="Acronycta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B912EAC3FFE6FF8A333CFDDDFCD7473D" box="[615,809,528,551]" italics="true" pageId="5" pageNumber="400">Acronycta bicolor</emphasis>
</taxonomicName>
/ [18]87, [
<collectingCountry id="F3717641FFE6FF8A32C5FDDDFC22473D" box="[926,988,528,551]" name="India" pageId="5" pageNumber="400">India</collectingCountry>
], Solun, May (coll. 
<collectionCode id="ED77AE14FFE6FF8A35EDFDDDFB0F473D" box="[1206,1265,528,551]" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="5" pageNumber="400">ZISP</collectionCode>
), genitalia slide Matov0396; 
<specimenCount id="9D60FD58FFE6FF8A3078FDFFFE784753" box="[291,390,562,585]" pageId="5" pageNumber="400" type="female">1 female</specimenCount>
, Tibet, Dsagar-Bg., 
<quantity id="4C9E9B34FFE6FF8A333CFDFFFD434753" box="[615,701,562,585]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="400" unit="m" value="3000.0">3000 m</quantity>
, Juli, Staatsslg. München, e. coll. F. Papp, genitalia slide AV1460 Volynkin (coll. 
<collectionCode id="ED77AE14FFE6FF8A3066FD98FE864776" box="[317,376,597,620]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="5" pageNumber="400">ZSM</collectionCode>
); 
<specimenCount id="9D60FD58FFE6FF8A30D0FD98FE1F4776" box="[395,481,597,620]" pageId="5" pageNumber="400" type="male">4 males</specimenCount>
, 
<specimenCount id="9D60FD58FFE6FF8A30B0FD98FDA84776" box="[491,598,597,620]" pageId="5" pageNumber="400" type="female">5 females</specimenCount>
, SW 
<collectingCountry id="F3717641FFE6FF8A33D4FD98FD2F4776" box="[655,721,597,620]" name="China" pageId="5" pageNumber="400">China</collectingCountry>
, Yunnan prov., Lijiang, 
<quantity id="4C9E9B34FFE6FF8A3289FD98FB9C4776" box="[978,1122,597,620]" metricMagnitude="3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="5" pageNumber="400" unit="m" value="3000.0" valueMax="4000.0" valueMin="2000.0">2000–4000m</quantity>
, 
<date id="FFD81011FFE6FF8A3537FD98FB264776" box="[1132,1240,597,620]" pageId="5" pageNumber="400" value="1934-05-18">18.v.1934</date>
; 
<date id="FFD81011FFE6FF8A35BFFD98FAF44776" box="[1252,1290,597,620]" pageId="5" pageNumber="400">9.v.</date>
, 
<date id="FFD81011FFE6FF8A3454FD98FABA4776" box="[1295,1348,597,620]" pageId="5" pageNumber="400">10.v.</date>
, 
<date id="FFD81011FFE6FF8A3412FD98FA804776" box="[1353,1406,597,620]" pageId="5" pageNumber="400">12.v.</date>
(
<specimenCount id="9D60FD58FFE6FF8A34D6FD98FF104795" pageId="5" pageNumber="400" type="female">2 females</specimenCount>
), 
<date id="FFD81011FFE6FF8A3058FDB5FEC64795" box="[259,312,632,655]" pageId="5" pageNumber="400" value="1935-05">13.v.</date>
, 
<date id="FFD81011FFE6FF8A301CFDB5FE834795" box="[327,381,632,655]" pageId="5" pageNumber="400" value="1935-05">24.v.</date>
, 
<date id="FFD81011FFE6FF8A30D0FDB5FE2F4795" box="[395,465,632,655]" pageId="5" pageNumber="400" value="1935-07">23.vii.</date>
, 
<date id="FFD81011FFE6FF8A3084FDB5FD9E4795" box="[479,608,632,655]" pageId="5" pageNumber="400" value="1935-08-22">22.viii.1935</date>
(coll. 
<collectionCode id="ED77AE14FFE6FF8A33F0FDB5FD074795" LSID="urn:lsid:biocol.org:col:34613" box="[683,761,632,655]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="5" pageNumber="400">ZFMK</collectionCode>
), genitalia slides Stüning 2288-DS, Kononenko 1780- 
<collectionCode id="ED77AE14FFE6FF8A3412FDB5FA664795" LSID="urn:lsid:biocol.org:col:34613" box="[1353,1432,632,655]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="5" pageNumber="400">ZFMK</collectionCode>
, 1781- 
<collectionCode id="ED77AE14FFE6FF8A3189FD57FEDC47AB" LSID="urn:lsid:biocol.org:col:34613" box="[210,290,666,689]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="5" pageNumber="400">ZFMK</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C37C655AFFE6FF8A319CFD70FAF04645" pageId="5" pageNumber="400" type="diagnosis">
<paragraph id="8BD936D1FFE6FF8A319CFD70FAF04645" blockId="5.[151,1436,354,897]" pageId="5" pageNumber="400">
<emphasis id="B912EAC3FFE6FF8A319CFD70FECA47CE" bold="true" box="[199,308,701,724]" pageId="5" pageNumber="400">Diagnosis</emphasis>
. 
<taxonomicName id="4C664D52FFE6FF8A3019FD70FE0C47CE" box="[322,498,701,724]" class="Insecta" family="Noctuidae" genus="Acronicta" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="400" phylum="Arthropoda" rank="subSpecies" species="grumi" subSpecies="bicolor">
<emphasis id="B912EAC3FFE6FF8A3019FD70FE0C47CE" box="[322,498,701,724]" italics="true" pageId="5" pageNumber="400">A. grumi bicolor</emphasis>
</taxonomicName>
(
<figureCitation id="135D2A54FFE6FF8A335BFD70FD9E47CE" box="[512,608,701,724]" captionStart="FIGURES 1 – 10" captionStartId="2.[151,258,1860,1882]" captionTargetBox="[157,1436,198,1838]" captionTargetId="figure@2.[151,1436,191,1839]" captionTargetPageId="2" captionText="FIGURES 1 – 10. Acronicta spp.: adults. 1, A. grumi, holotype male, Tien Shan (ZISP); 2, A. grumi, male, holotype of A. tiena, [Kyrgyzstan, Kirgizsky Ridge] &quot; Alexandergebirge &quot; (NKMB, photo by W. Mey); 3, A. grumi, female, [Kyrgyzstan, Kirgizsky Ridge] &quot; Alexandergeb. &quot; (ZSM); 4, A. grumi bicolor, syntype of Acronycta bicolor, female, N India, Solun (© NHM) (photo by A. Zilli); 5, A. grumi bicolor, female, N India, Solun (ZISP); 6, A. grumi bicolor, female, China, Tibet (ZSM); 7, A. rumicis, female, West Siberia (AVB); 8, A. rumicis, male, SW Mongolia (AVB); 9, A. indica, male, N India, Sikkim (ZISP); 10, A. megacephala, type species of Subacronicta, European part of Russia, Tula reg. (ZISP)." httpUri="https://zenodo.org/record/245569/files/figure.png" pageId="5" pageNumber="400">Figs 4–6</figureCitation>
) differs externally from the nominate subspecies (
<figureCitation id="135D2A54FFE6FF8A352EFD70FB2D47CE" box="[1141,1235,701,724]" captionStart="FIGURES 1 – 10" captionStartId="2.[151,258,1860,1882]" captionTargetBox="[157,1436,198,1838]" captionTargetId="figure@2.[151,1436,191,1839]" captionTargetPageId="2" captionText="FIGURES 1 – 10. Acronicta spp.: adults. 1, A. grumi, holotype male, Tien Shan (ZISP); 2, A. grumi, male, holotype of A. tiena, [Kyrgyzstan, Kirgizsky Ridge] &quot; Alexandergebirge &quot; (NKMB, photo by W. Mey); 3, A. grumi, female, [Kyrgyzstan, Kirgizsky Ridge] &quot; Alexandergeb. &quot; (ZSM); 4, A. grumi bicolor, syntype of Acronycta bicolor, female, N India, Solun (© NHM) (photo by A. Zilli); 5, A. grumi bicolor, female, N India, Solun (ZISP); 6, A. grumi bicolor, female, China, Tibet (ZSM); 7, A. rumicis, female, West Siberia (AVB); 8, A. rumicis, male, SW Mongolia (AVB); 9, A. indica, male, N India, Sikkim (ZISP); 10, A. megacephala, type species of Subacronicta, European part of Russia, Tula reg. (ZISP)." httpUri="https://zenodo.org/record/245569/files/figure.png" pageId="5" pageNumber="400">Figs 1–3</figureCitation>
) by the darker and more monotonous blackish ground colour of forewings; in the male genitalia, 
<taxonomicName id="4C664D52FFE6FF8A328CFD2DFB7647ED" box="[983,1160,736,759]" class="Insecta" family="Noctuidae" genus="Acronicta" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="400" phylum="Arthropoda" rank="subSpecies" species="grumi" subSpecies="bicolor">
<emphasis id="B912EAC3FFE6FF8A328CFD2DFB7647ED" box="[983,1160,736,759]" italics="true" pageId="5" pageNumber="400">A. grumi bicolor</emphasis>
</taxonomicName>
(
<figureCitation id="135D2A54FFE6FF8A35CCFD2DFB1A47ED" box="[1175,1252,736,759]" captionStart="FIGURES 11 – 16" captionStartId="3.[151,258,1951,1973]" captionTargetBox="[187,1431,198,1929]" captionTargetId="figure@3.[151,1436,191,1930]" captionTargetPageId="3" captionText="FIGURES 11 – 16. Acronicta spp.: male genitalia. 11, A. grumi, holotype, NW China, slide Matov 0144; 12, A. grumi, holotype of A. tiena, Kyrgyzstan, slide MB 358 Boursin (photo by W. Mey); 13, A. grumi bicolor, China, slide Stüning 2288 - DS; 14, A. rumicis, East Kazakhstan, slide AV 0931 Volynkin; 15, A. indica, N India, slide Matov 0385; 16, A. digna, Russian Far East, slide AV 1499 Volynkin." httpUri="https://zenodo.org/record/245570/files/figure.png" pageId="5" pageNumber="400">Fig. 13</figureCitation>
) has a somewhat narrower apical part of valva, more or less C-shaped ampulla (in the nominate subspecies the ampulla is more or less Sshaped), less sclerotized medial part of vesica, and a longer and narrower sclerotized plate in medial part of vesica; in the female genitalia, 
<taxonomicName id="4C664D52FFE6FF8A3014FC85FE014645" box="[335,511,840,863]" class="Insecta" family="Noctuidae" genus="Acronicta" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="400" phylum="Arthropoda" rank="subSpecies" species="grumi" subSpecies="bicolor">
<emphasis id="B912EAC3FFE6FF8A3014FC85FE014645" box="[335,511,840,863]" italics="true" pageId="5" pageNumber="400">A. grumi bicolor</emphasis>
</taxonomicName>
(
<figureCitation id="135D2A54FFE6FF8A3355FC85FD764645" box="[526,648,840,863]" captionStart="FIGURES 17 – 24" captionStartId="4.[151,258,1637,1659]" captionTargetBox="[155,1431,200,1614]" captionTargetId="figure@4.[151,1436,191,1616]" captionTargetPageId="4" captionText="FIGURES 17 – 24. Acronicta spp.: male (17 – 19) and female (20 – 24) genitalia. 17, A. megacephala, type species of Subacronicta, Austria, Vienna, slide AV 1024 Volynkin; 18, A. grumi, aedeagus of holotype, diagnostically important sclerotized elements of vesica; 19, A. grumi, aedeagus of tiena lectotype, diagnostically important sclerotized elements of vesica; 20, A. grumi, Alexandergebirge, slide AV 1459 Volynkin; 21, A. grumi bicolor, China, slide AV 1460 Volynkin; 22, A. grumi bicolor, N India, Solun, slide Matov 0396; 23, A. rumicis, Russia, Altai Mts., slide AV 1501 Volynkin; 24, A. indica, Nepal, slide Stüning 2289 - DS." httpUri="https://zenodo.org/record/245571/files/figure.png" pageId="5" pageNumber="400">Figs 21, 22</figureCitation>
) has a more chalice-shaped and stronger sclerotized antrum.
</paragraph>
</subSubSection>
<subSubSection id="C37C655AFFE6FF8A319CFCA7FBB9469B" box="[199,1095,874,897]" pageId="5" pageNumber="400" type="distribution">
<paragraph id="8BD936D1FFE6FF8A319CFCA7FBB9469B" blockId="5.[151,1436,354,897]" box="[199,1095,874,897]" pageId="5" pageNumber="400">
<emphasis id="B912EAC3FFE6FF8A319CFCA7FEAC469B" bold="true" box="[199,338,874,897]" pageId="5" pageNumber="400">Distribution</emphasis>
. The subspecies is known from the Himalaya and southwestern 
<collectingCountry id="F3717641FFE6FF8A355BFCA7FBBD469B" box="[1024,1091,874,897]" name="China" pageId="5" pageNumber="400">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>