A Review of the Australian Species of Thevenetimyia Bigot, 1875 (Bombyliidae, Bombyliinae, Eclimini), with Description of Four New Species and the Pupal Case of T. longipalpis (Hardy) Author Li, Xuankun Author Rodrigues, Paula Fernanda Motta Author Lamas, Carlos José Einicker Author Yeates, David K. text Records of the Australian Museum 2018 2018-05-30 70 3 331 375 http://dx.doi.org/10.3853/j.2201-4349.70.2018.1678 journal article 10.3853/j.2201-4349.70.2018.1678 2201-4349 5237395 4D83F929-371B-4BFF-A271-54CE7B13C5A1 Thevenetimyia nigrapicalis ( Roberts, 1929 ) Figs 19–22 Eclimus nigrapicalis Roberts, 1929: 576 . Type-locality: Australia (QLD); holotype , QM ( Fig. 19 ). Thevenetimyia nigrapicalis ( Roberts, 1929 ) .– Hall, 1969: 65 . Types and other specimens examined . Holotype ♂ , Australia , QLD , Goondiwindi , 29 Dec 1927 , F. Roberts , QM D3525 . Paratypes : allotype ♀ , same data as holotype, QM T6499 ; 1♀ , QLD, Goondiwindi , 20 Dec 1927 , GB , QM T183310 /D.520; 3♂♂ , QLD, Goondiwindi , Jan 1928 , G. R. Bass , QM T183307 / D.523, QM T183308 / D.524, QM T183309 /D.525; 2♂♂ , QLD, Goondiwindi , Dec 1927 , G. R. Bass , QM T183305 / D.522, QM T183306 /D.521; 1♂ , Eidsvold , Dec 29– Apr 30, T. L. Bancroft , ANIC 29- 041298 . NSW , 1♂ , Mt Boppy , nr Cobar , 25 Nov 1949 , S. J. Paramonov , ANIC 29-008908 ; 1♂ , NSW, Mt Boppy , nr Cobar , 25 Nov 1949 , S. J. Paramonov , ANIC 29-008908 ; 1♂ , Carnarvon Nat. Park , Mt Moffatt Section , summit of Mt Moffatt , 1097 m , 25°04'S 148°03'E , 25 Nov 1995 , D. K. Yeates , C. J. Burwell , QM ; 1♂ , Mt Moffat NP, Mt Moffat summit, 15 Dec 1987 , D. K. Yeates , QM ; 1♀ , Basalt Ck , 4 km E Mitchell , 23 Nov 1979 , K. L. Walker , on Eucalyptus populnea F. Muell., QM ; 1♀ , Westwood , 20 Oct 1927 , A . P. Dodd , NHMUK ; 1♂ , Goondiwindi , 29 Dec 1927 , FH Roberts , NHMUK . NT , 1♂ , Border Waterhole , 15 km W Musselbrook Resource Centre , Lawn Hill NP, 200 m , NT 18°36'44"S 137°59'30"E , 19 Apr 1995 , G. Daniels , M. A . Schneider , QM . Figure 19. Thevenetimyia nigrapicalis ( Roberts, 1929 ) ♂ holotype: (a) dorsal; (b) lateral; (c) wing. Scale bars = 1 mm. Figure 20. Thevenetimyia nigrapicalis ( Roberts, 1929 ) ♂: (a) dorsal; (b) lateral; (c) flagellum; (d) head, dorsal; (e) head, lateral; (f) head, frontal; (g) head, profile. Scale bars = 1 mm (a, b, d–h); = 0.1 mm (c). Figure 21. Thevenetimyia nigrapicalis ( Roberts, 1929 ) ♀ allotype: (a) dorsal; (b) lateral; (c) head, dorsal; (d) head, lateral; (e) wing; (f) head, frontal; (g) head, profile. Scale bars = 1 mm. Figure 22. Thevenetimyia nigrapicalis ( Roberts, 1929 ) ♂ genitalia: (a) genital capsule, dorsal; (b) genital capsule, ventral; (c) genital capsule, lateral; (d) epandrium, dorsal; (e) epandrium, lateral. Scale bars = 0.1 mm. Diagnosis . Medium-sized Thevenetimyia , wing membrane mostly hyaline, dark infuscation present on middle part of anterior half; costa with two rows of small spines; scutum with two rows of distinct short, thick spines on pruinescence stripes, three uneven rows and some adjacent, separate spines; lateral areas of abdominal tergites 6 and 7 with dense long black hairs, lateral areas of tergite 8 dense long white hairs; outer apex of gonocoxite relatively narrow. Redescription . Male . Body length 9.5–10.3 mm , wing length 8.1–8.7 mm . Head. Head about 1.6× wider than long, mostly blackish with thick pale pruinescence and covered in admixed white to blackish brown hairs. Eye narrowly connected, by 0.5× length of ocellar tubercle. Frons slender and long, upper triangular section small, lower triangular section black with thick pale pruinescence, middle narrow part about 0.6× length of frons. Ocellar tubercle slightly raised, brown to blackish brown with grey pruinescence, with blackish brown hairs. Face with thick pale pruinescence and admixed dark yellow and white hairs, parafacial area dark brown and bare. Gena with thick pale pruinescence and long white hairs. Clypeus swollen, with thick pale pruinescence and white and admixed some dark yellow hairs. Occiput with long white hairs except dorsal margin admixed with brown to black hairs. Posterior eye margin slightly sinuous. Antennae blackish brown; scape and pedicel with thin pale pruinescence and black hairs, ventral face of scape admixed with white hairs. Scape 3.1× as long as wide, and 2.9× as long as pedicel, uniform from base to apex. Pedicel 1.2× as long as wide. Flagellum 5.8× as long as wide, 1.3× as long as scape+pedicel (1.8× as long as scape), conical and slightly laterally compressed, one-segmented with apical style ( Fig. 20c ). Palp thin and long, extending beyond oral cavity, black with blackish brown hairs, two-segmented, with palpal pit. Mouthparts slender, 4.0× as long as eye length (2.4× as long as head length), labella thin and filiform ( Fig. 20e ). Thorax. Integumental colour of scutum and scutellum mostly black with sparse grey pruinescence, scutum with two pruinescence stripes, and pruinescence more dense around margins. Scutum and scutellum with admixed white and pale yellow hairs and long pale yellow scales. Three notopleural setae present. Postalar callus and posterior margin of scutellum with strong hairs. Notopleural setae and strong hairs on postalar callus pale yellow, setae on posterior margin of scutellum pale yellow to black. Scutum with two rows of distinct short, thick spines on pruinescence stripes, three uneven rows and some adjacent, separate spines. Pleura and coxae black with thick grey pruinescence, anepisternum, katepisternum and metepisternum with long white hairs, mediotergite with dense white hairs and setae, anepimeron, meron and laterotergite bare. Legs. Legs blackish brown, mostly covered with white scales, except hind tibia with blackish brown scales. Fore and mid femora with long white hairs on ventral face, hind femur with a row of short ventral bristles. Other hairs and bristles on legs short and blackish brown to black. Fore tibia 2.2× longer than fore basitarsus, mid tibia 2.0× longer than mid basitarsus, hind tibia 1.9× longer than hind basitarsus. Wings. Wing membrane mostly hyaline, dark infuscation present on middle part of anterior half, proximal side covering crossvein r-m , distal side reaching to apex of vein R 1 , without a distinct margin. Cell r 5 widely open; cell br much longer than cell bm , crossvein r-m arising a little over half way from the base of cell dm ; crossvein m-m long, and 2.0× as long as crossvein r-m ; cell cup open ( Fig. 19c ). Costa with two rows of short, thick spines. Haltere stem pale yellow, knob black. Abdomen. Integumental colour of tergites black with thin pale pruinescence, except lateral margin, posterior half of tergite 7 and tergites 8 and 9 with thick pruinescence. Tergites mostly covered with black scales and short black hairs. Tergite 1 covered with long white hairs. Tergites 2 to 5 with long white hairs admixed with black scales, posterior and lateral margins with white scales. Tergites 6 and 7 with white scales on posterior half, and lateral margins with dense long black hairs. Tergite 8 with dense long white hairs laterally. Genitalia. Epandrium wide and short, posterior margin slightly convex ( Fig. 22d ). Hypandrium present. Gonocoxal apex slightly narrower than the base in dorsal view, ejaculatory apodeme large ( Figs 22a, b ); gonocoxal apodeme strong and incurved; lateral ejaculatory process large, 2.5× longer than wide, apex not expanded; inner apex of gonocoxite rounded, elongate and broad; outer apex of gonocoxite relatively narrow; dorsal bridge without lateral hollow ( Fig. 22c ); gonostylus slender and pointed dorsally, 3.4× longer than wide. Female . Body length 10.0 mm, wing length 7.5 mm . Very similar to male, except frons 2.5× as wide as ocellar tubercle, with short black hairs on dorsal half ( Fig. 21c ). Wing completely hyaline, costa without small spines ( Fig. 21e ). Abdominal tergite 8 devoid silver hairs. Remarks . Thevenetimyia nigrapicalis ( Roberts, 1929 ) is redescribed and images of external and internal characters are provided. Distribution . NSW, QLD.