Re-description of Grammapsychops lebedevi Martynova, 1954 (Neuroptera: Psychopsidae) with notes on the Late Cretaceous psychopsoids
Author
Makarkin, Vladimir N.
text
Zootaxa
2018
2018-11-26
4524
5
581
594
journal article
27908
10.11646/zootaxa.4524.5.5
fcefbd6c-a129-4576-91c8-569c8e0e37f6
1175-5326
2610783
297DB9B9-F1F8-46C0-8E43-95EBA0906B9F
Genus
Grammapsychops
Martynova, 1954
Grammapsychops
Martynova, 1954
: 1167
, 1169
[
Psychopsididae
(sic)];
Carpenter 1992
: 351
[
Psychopsidae
];
Oswald 1993b
: 48
[
Psychopsidae
];
Andersen 2001
: 435
[
Psychopsidae
s.l.: Group IV];
Makarkin & Archibald 2003
: 177
[
Psychopsidae
?];
Jepson
et al
. 2009
: 1330
[
Osmylopsychopidae
];
Peng
et al
. 2011
: 218
, 221, 225 [
Psychopsidae
];
Makarkin
et al
. 2012
: 59
[
Psychopsidae
];
Makarkin & Khramov 2015
: 411
[
Osmylopsychopidae
?];
Peng
et al
. 2016
: 262
[
Osmylopsychopidae
?].
Grammopsychops
[sic, misspelled]:
Martynova 1962
: 278 [
Psychopidae
(sic)];
Martins-Neto 1997
: 70, 71 [
Psychopsidae
];
Martins-Neto & Rodrigues 2010
: 6 [
Psychopsidae
].
Type
species.
Grammapsychops lebedevi
Martynova, 1954
, by original designation.
Diagnosis
. In hind wing, costal space extremely broad, with numerous crossveins arranged mainly in several gradate series; ScP and RA connected distally by relatively long crossvein; R then basal part of RP very stout; posterior trace of RA with three long distal anteriorly-directed branches; RA space very narrow; branches of RP deeply dichotomously branched; RP1–RP6 stout proximally; RP1 shallowly forked; M forked basally, only slightly distad origin of RP1; CuA strongly pectinately branched, with very oblique branches; CuP with three long, very oblique branches; many crossveins in radial to cubital spaces arranged mainly in irregular gradate series.
Species included
.
Type
species only.
Comments
. The venation of the hind wing of
Grammapsychops
is most similar to those of two Cretaceous genera, i.e.,
Pulchroptilonia
Martins-Neto, 1997
and
Embaneura
Zalessky, 1953
.
Zalessky (1953)
interpreted the only known wing of
E. vachrameevi
as a forewing, but judging from its venation this is most probably a hind wing. Unfortunately, this cannot be verified due to the current depository of the
holotype
of
E. vachrameevi
Zalessky, 1953
being unknown.
Grammapsychops
easily differs from these genera by the presence of numerous crossveins in the costal space of the hind wing (no crossveins are present in
Pulchroptilonia
and
Embaneura
).
The hind wing of
Grammapsychops
is also similar to that of
Alloepipsychopsis
Makarkin
et al
., 2012
from the Early Cretaceous (late Barremian) Yixian Formation, but the latter has its RP1 deeply forked.
The hind wing venation of other genera of
Psychopsidae
(and
Osmylopsychopidae
) is dissimilar to that of
Grammapsychops
. Unfortunately, however, the hind wings are not known in all fossil genera of these families.