A new Helluodes Westwood species from Western Ghats with notes on habitat, distribution and phylogenetic relationships of genera Helluodes Westwood and Physocrotaphus Parry of the tribe Physocrotaphini Chaudoir (Insecta: Coleoptera: Carabidae: Anthiinae) Author Sabu, Thomas K. Litter Entomology Research Unit, P. G. & Research Department of Zoology, St. Joseph's College, Devagiri, Calicut, Kerala, 673008, India Department of Entomology, College of Natural Resources & Environment, South China Agricultural University, Wushan, Guangzhou, 510640, China Corresponding author. E-mail: sabukthomas @ gmail. com sabukthomas@gmail.com Author Abhitha, Prabhakar Litter Entomology Research Unit, P. G. & Research Department of Zoology, St. Joseph's College, Devagiri, Calicut, Kerala, 673008, India Department of Entomology, College of Natural Resources & Environment, South China Agricultural University, Wushan, Guangzhou, 510640, China Corresponding author. E-mail: sabukthomas @ gmail. com Author Zhao, Danyang text Zootaxa 2008 2008-04-09 1745 1 30 46 https://biotaxa.org/Zootaxa/article/view/zootaxa.1745.1.3 journal article 10.11646/zootaxa.1745.1.3 1175­5334 5108342 Key to the species of Helluodes Westwood 1. Anterior margin of labrum with a distinct median emargination between rounded anterior angles ( Fig. 12 ); anterior angles of pronotum expanded, rounded and weakly acute; pronotum distinctly sinuated anterior to posterior angles; posterior angles of pronotum strongly reflexed and sharp at apex ( Fig. 14 ); ligula slen- der, widest at middle; TLA = 25–27.5 .................................................................... H. westwoodi Chaudoir. - Anterior margin of labrum nearly straight or slightly emarginated between rounded anterior angles; anterior angles of pronotum expanded and rounded; pronotum weakly sinuated anterior to posterior angles; posterior angles of pronotum weakly reflexed and blunt at apex ( Figs. 8, 11 ); ligula slender and narrower towards apex................................................................................................................................................2 2 Anterior margin of labrum distinctly crenulate with 4 distinct indentations between anterior angles ( Fig. 6 ); maxillary palpomere 4 truncate and broadly dilated from middle to apex ( Fig. 7 ); TLA = 24–26.2 ...... .................................................................................................................................. H. devagiriensis n.sp. - Anterior margin of labrum without crenulation and slightly emarginate between anterior angles ( Fig. 9 ); maxillary palpomere 4 roundly narrowed at apex and roundly dilated from middle to apex ( Fig. 10 ); TLA = 24.5 .................................................................................................................. H.taprobanae Westwood.