Notes on Rhithrodytes BAMEUL, 1989, with the description of R. minimus nov. sp. from Algeria (Coleoptera, Dytiscidae. Hydroporini, Siettitiina)
Author
Fery, Hans
text
Linzer biologische Beiträge
2016
2016-07-30
48
1
431
450
journal article
10.5281/zenodo.5416650
0253-116X
5416650
Rhithrodytes argaensis
BILTON & FERY
,
1996
(new status)
Rhithrodytes agnus
argaensis
BILTON & FERY, 1996: 919
;
RIBERA et al. 1999: 60
;
RIBERA 2003: 477
; FERY & FRESNEDA 2007: 138;
ABELLÁN et al. 2013: 123
and Appendices (phylogeny).
Ty pe locality: "
Portugal
, Minho, NE
Viana do Castelo
, Serra de Arga, stream at
700 m
".
For the description of
R. argaensis
and comparison with
Rhithrodytes agnus
FOSTER, 1992
see BILTON & FERY (1996). The
holotype
is stored in the NMW. The habitus of this species is given in
Fig. 16
, the male genitalia in
Figs 29 and 33
; see
Figs 15
,
28 and 32
for
R. agnus
.
So far this taxon was treated as a subspecies of
R. agnus
. Because the
type
localities of the two subspecies are only about
10 km
away from each other, the authors of the original description hesitated to assign specific rank to
R. argaensis
. On the other hand, they found that the two taxa are obviously incapable of dispersal by flight (BILTON & FERY 1996: 924). This is why the authors suspected that both subspecies might be given specific rank following (future) genetic investigations and wrote "... but it would appear that these represent taxa on their way to speciation." (BILTON & FERY 1996: 928).
Recently, such genetic investigations have been presented by
ABELLÁN et al. (2013
, appendices) and Ignacio Ribera (Barcelona,
Spain
) communicated privately that if both subspecies have "... enough morphological differences they can be upgraded to species... genetically they are very close, but well separated." In particular, the differences in the male genitalia are very distinct (see
Figs 28 and 29
), and thus the rank of
argaensis
is here elevated from subspecific to specific.
The data of the specimens used for the genetic studies in
ABELLÁN et al. (2013)
are: "
28.5.2006
(P)
Viana do Castelo
, ca.
6 km
N
Ponte de Lima
,
W Labruja
, rest ponds of brook under bridge, UTM ca. NG 324311,
H. Fery
;
COI:
HF931143
;
16S:
HF931362
" and "
9/v/2005
Serra de Arga
, pools on summit,
D.T. Bilton
; COI:
HF931183
, 16S:
HF931405
"
.