A Taxonomic Revision of Nearctic Conostigmus (Hymenoptera: Ceraphronoidea: Megaspilidae)
Author
Trietsch, Carolyn
Author
Mikó, István
Author
Ezray, Briana
Author
Deans, Andrew R.
text
Zootaxa
2020
2020-06-15
4792
1
1
155
journal article
21681
10.11646/zootaxa.4792.1.1
dd8ef826-0c0f-4261-b127-1d1afa7f5601
1175-5326
3895976
326F6A15-216E-439A-AD59-3CDF7551D3F6
Dendrocerus arietinus
(Provancher, 1887)
,
new combination
Figs. 49
,
50
Species Comments and History.
Isostasis arietinus
Provancher, 1887
was described from a single female specimen. Provancher named the species
arietinus
or “tête de belier” after the curve of the antennae, which he believed resembled the horns of a ram. Provancher (1888) transferred the species from
Isostasis
to the genus
Baeoneura
, another genus described by
Förster (1856)
.
Harrington (1900)
transferred the species from Platygastroidea to
Ceraphronoidea
, remarking that the description “answers very well to
Megaspilus Harringtoni
, Ashm.
”, and
Kieffer (1914)
subsequently redescribed the species and transferred it to
Conostigmus
. Gahan and Rohwer (1917) designated a
Baeoneura arietina
specimen as a
lectotype
at the ULCQ, and noted that the specimen was badly glued.
Masner (1969)
echoed this statement on the condition of the specimen and reported that the specimen was a
Conostigmus
species.
Provancher (1887) originally described the species
arietinus
as a member of the genus “
Isostasis
”, which Provancher attributed to Förster; however, Förster never described a genus “
Isostasis
”. Instead, Förster had described the platygastrid genus
Isostasius
(
Hymenoptera
,
Platygastridae
) (
Förster, 1856
).
Dessart (1996)
later designated
Isostasis
Provancher, 1888
not as an incorrect spelling but as a valid genus that was synonymous with
Conostigmus
Dahlbom, 1858
.
Johnson and Musetti (2004)
do not accept Dessart’s action and instead consider
Isostasis
as a misspelling of Förster’s
Isostasius
, pointing out that such an action would set a precedent leading “to the adoption of untold numbers of junior homonyms!” (pg. 6).
We mention the troubled history of this species to comment that Dessart did not directly observe the
holotype
, but considered the species a
Conostigmus
based on the assertions of
Kieffer (1914)
and
Masner (1969)
. Upon examining the female
lectotype
specimen of
Conostigmus arietinus
(Provancher), 1887
at the ULQC, however, we realized that this specimen was a
Dendrocerus
based on its close resemblance to the female
holotype
of
D. penmaricus
(see
Figs. 49
,
50
for comparison with the female
holotype
of
Dendrocerus penmaricus
(Ashmead), 1893
deposited in the USNM). The species also has the following combination of
Dendrocerus
-like characters: ocelli in an obtuse triangle (POL greater than LOL), where the two posterior ocelli are closer to the compound eyes than to each other (POL greater than OOL); mesometapleural sulcus arched; and facial pit absent.
This species likely belongs to the
D. penmaricus
species group, and may even be the same species as
D. penmaricus
, as the female type of
arietinus
shares the following characters with the female
holotype
of
D. penmaricus
: ocelli in an obtuse triangle; facial pit absent; median process on the intertorular carina present and blunt; preoccipital furrow present and the anterior end reaching the anterior ocellus; postocellar furrow absent; mandibles with two teeth; posterior end of the notaulus not adjacent to the median mesoscutal sulcus; and the general habitus (in particular, the shape of the head and the size of the eyes in relation to the head in lateral view are unique). The type specimen of
arietinus
is so poorly glued that it is not possible to confirm the absence of a sternaulus, which would be another indicator that this species is indeed a
Dendrocerus
.
Because the female
type
is in such poor condition and the male of the species is unknown, it is not possible to confirm whether
D. arietinus
is synonymous with
D. penmaricus
. For now, we consider
D. arietinus
and
D. penmaricus
as separate species, but we move
D. arietinus
from
Conostigmus
to
Dendrocerus
and consider
D. arietinus
as a member of the
D. penmaricus
species group.
Material Examined.
Lectotype
female:
CANADA
:
PSUC
_
FEM 148706
(
ULQC
).