A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae) Author Pimvichai, Piyatida 4D882A89-E0C1-4F78-93E7-FA1B5A2650F2 Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai District, Maha Sarakham 44150, Thailand. & urn: lsid: zoobank. org: author: 4 D 882 A 89 - E 0 C 1 - 4 F 78 - 93 E 7 - FA 1 B 5 A 2650 F 2 & Corresponding author: piyatida _ pimvichai @ yahoo. com piyatida_pimvichai@yahoo.com Author Enghoff, Henrik FB09A817-000D-43C3-BCC4-2BC1E5373635 Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, DK- 2100 Copenhagen Ø, Denmark. & Email: henghoff @ snm. ku. dk & urn: lsid: zoobank. org: author: FB 09 A 817 - 000 D- 43 C 3 - BCC 4 - 2 BC 1 E 5373635 henghoff@snm.ku.dk Author Panha, Somsak AC935098-D901-4F35-A414-4B0D4FE44E79 Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand. & Email: somsakp @ sc. chula. ac. th & urn: lsid: zoobank. org: author: AC 935098 - D 901 - 4 F 35 - A 414 - 4 B 0 D 4 FE 44 E 79 somsakp@sc.chula.ac.th Author Backeljau, Thierry E6F42575-36AE-4AD7-98C6-D083EF052568 University of Antwerp, Evolutionary Ecology Group, Groenenborgerlaan 171, B- 2020 Antwerp, Belgium and Royal Belgian Institute of Natural Sciences, OD Taxonomy and Phylogeny, and JEMU, Vautierstraat 29, B- 1000 Brussels, Belgium. & Email: Thierry. Backeljau @ naturalsciences. be & urn: lsid: zoobank. org: author: E 6 F 42575 - 36 AE- 4 AD 7 - 98 C 6 - D 083 EF 052568 ackeljau@naturalsciences.be text European Journal of Taxonomy 2016 2016-05-24 199 1 37 journal article 21904 10.5852/ejt.2016.199 d88184f4-ceef-4392-a444-0ac1552778d3 2118-9773 3840484 EE35A675-06C7-4FF9-9EE0-0854D95036C4 Thyropygus navychula sp. nov. urn:lsid:zoobank.org:act: 01AC06C9-EEF0-4211-9837-28FE2C12C68E Figs 6 A–D, 11A Diagnosis A species of the opinatus subgroup. Mesal margin of anterior coxal fold ( alp ) with fine serrations. Similar in this respect to T. opinatus . Differs from this species by having the lateral process ( alp ) Fig. 6. Thyropygus navychula sp. nov. , from Surin Islands, holotype (CUMZ-D00095), ³, gonopods. A . Anterior view, left telopodite removed. B . Posterior view, left telopodite removed. C . Left telopodite, posterior-mesal view. D . Left telopodite, anterior-lateral view. slender, regularly curved and by having the mesal process ( amp ) broadly expanded and forming a strong longitudinal crest ( lc ) in posterior view. Etymology The species is named after the Royal Thai Navy, in recognition of their kind assistance which enabled us to pursue the necessary fieldwork at the type locality, and also after Chulalongkorn University where pink is the symbolic colour of the university, indirectly referring to the pink legs of the species; the species name is treated as a noun in apposition. Material examined Holotype THAILAND : ³, Phang-Nga Province , Khuraburi District , Surin Islands , 9°24'20" N , 97°51'38" E , 7 Apr. 2012 , leg. S. Panha , P. Pimvichai and members of the Animal Systematics Research Unit ( CUMZ-D00095 ). Paratypes THAILAND : 5 ³³, 1 ♀ , same data as holotype ( CUMZ-D 00089); 2 ³³, same data as holotype ( ZMUC ). Description Adult males with 58–62 podous rings, no apodous rings. Length 11–12 cm , width 6.2–7.2 mm . Adult female with 60 podous rings, no apodous rings. Length 14 cm , width 8.0 mm. Overall colour of living animal ( Fig. 11A ) dark brown, shiny. Legs and antennae pink; preserved specimen with head, antennae, prozona, legs, epiproct, paraprocts and hypoproct brown; metazona dark brown. GONOPODS ( Fig. 6 A–D). Anterior coxal fold ( ac ; Fig. 6A ): lateral process ( alp ) slender, regularly curved, mesal margin with fine serrations; mesal process ( amp ) almost as long as alp , broadly expanded, apically sharp, straight distad, in posterior view mesal margin of amp forming strong longitudinal crest ( lc ). Posterior coxal fold ( pc ; Fig. 6B ) basally with moderately high lateral paracoxites ( px ), distally with two processes: mesal process ( pmp ) shorter than plp , curving behind plp ; lateral process ( plp ) flattened, directed distad. Telopodite ( Fig. 6 C–D) leaving coxite between pmp and plp ; femoral spine ( fe ) very long, curving almost exclusively in horizontal plane; tibial spine ( ti ) very long, curving in horizontal plane close to basal part of fe ; spatulate lobe ( sl ) terminating in sharp spine; palette ( pa ) simple, gutterlike, distally with about twelve brownish blepharochaetae ( bp ), laterally with few sharp spines. DNA barcode The GenBank accession number of the barcode of one of the paratype is KU306522 (voucher code CUMZ-D00089). Distribution ( Fig. 12 ) Known only from the type locality.