Phylogenetic status of some unique species representing Blapstinina Mulsant & Rey (Tenebrionidae: Blaptinae: Opatrini), and implications for continued study of the subtribe Author Kamiński, Marcin Jan 0000-0002-1286-950X Zoological Museum, Museum and Institute of Zoology, Polish Academy of Sciences, Wilcza 64, 00 - 679 Warszawa, Poland & Purdue University, Department of Entomology, 901 W. State Street, West Lafayette, IN 47907, USA https: // orcid. org / 0000 - 0002 - 1286 - 950 X Author Lumen, Ryan 0000-0002-3958-7596 Zoological Museum, Museum and Institute of Zoology, Polish Academy of Sciences, Wilcza 64, 00 - 679 Warszawa, Poland & https: // orcid. org / 0000 - 0002 - 3958 - 7596 Author Kanda, Kojun 0000-0001-5561-8471 USDA Systematic Entomology Laboratory, c / o Smithsonian Institution, National Museum of Natural History, Washington, DC, U. S. A. https: // orcid. org / 0000 - 0001 - 5561 - 8471 Author Smith, Aaron Dennis 0000-0002-1286-950X Purdue University, Department of Entomology, 901 W. State Street, West Lafayette, IN 47907, USA https: // orcid. org / 0000 - 0002 - 1286 - 950 X text Zootaxa 2022 2022-02-01 5093 5 519 532 journal article 20789 10.11646/zootaxa.5093.5.2 f349242c-a281-4590-aff8-0e52538be142 1175-5326 5964429 6D56585F-68D1-4AD0-83B6-87207480735B Lodinus araguae (Marcuzzi) , comb. et stat. nov. Austrocaribius venezuelensis araguae Marcuzzi, 1983: 259 Notes . Taking into consideration new morphological data concerning L. punctulatus and coarse differences in male genitalia structure (Figs 46 and 46 in Marcuzzi 1983 ; Figs 1 , 3D ) between the two previously recognized subspecies of L. venezuelensis (i.e., araguae and venezuelensis ), a decision is hereby made to elevate them to the species level. While describing this species Marcuzzi (1983) did not designate a holotype , nor explicitly state the number of studied specimens. As a result a lectotype designation is needed to fix the taxonomic status of the species. Studied material . LECTOTYPE (here designated) , male ( MIZ PAN ): “ Loma del Medio / San Sebastian / (Edo. Aragua )”, “BORDON leg. \ 3-VI-1962 ” ( Fig. 2E ) . PARALECTOTYPES , male ( MIZ PAN ): same data as lectotype; female ( MIZ PAN ): same data as lectotype with additional label: “ Paratypus 1986 \ Austrocaribius venezuelensis araguae ssp.\ Marcuzzi” ( Fig. 2F ) .