The genus Leucophenga (Diptera, Drosophilidae), part VI: the argentata species group from the East Asia, with morphological and molecular evidence Author Huang, Jia Author Chen, Hongwei text Zootaxa 2016 4161 2 207 227 journal article 10.11646/zootaxa.4161.2.4 45b6fbcc-533a-441f-b9db-ada4a02f10c7 1175-5326 264548 3B388468-4011-4EAB-A5E1-635DE1794536 Leucophenga quadricuspidata sp. nov. ( Figs. 2 E, 4G, 4H, 5M, 10) Diagnosis. This species is somewhat similar to L . fuscivena sp. nov. in the color patterns of abdominal tergites in male ( Fig. 5 G, H, M), but can be distinguished from the latter by having the aedeagus distolaterally with four acute processes each bearing a sensillum at tip ( Fig. 10 D). FIGURE 10. Leucophenga quadricuspidata sp. nov. , male terminalia in lateral view. A. Epandrium, surstylus and cercus; B. Hypandrium, paramere, gonopods and aedeagus; C. Paramere; D. Aedeagus and acute processes (ap). Scale bars = 0.1 mm. Description. Ocellar triangle brownish, with ca. 0–1 seta above ocellar setae. Frons yellowish to yellow. Pedicel yellowish to yellow; first flagellomere yellow. Clypeus brownish yellow. Palpus brownish. Mesonotum yellowish ( Fig. 4 G). Postpronotal lobe yellowish ( Fig. 4 H). Acrostichal setulae in ca. 6–8 irregular rows ( Fig. 4 G). Scutellum entirely yellowish ( Fig. 4 G). Pleura, katepisternum and meron yellowish ( Fig. 4 H). Wing ( Fig. 2 E): Costal vein between R2+3 and R4+5 distally with ca. 5–6 peg-like spinules on ventral surface. Only 3rd and 4th abdominal tergites with brownish to black patches in male ( Fig. 5 M). Male terminalia: Epandrium pubescent medially and near posterior margin, with ca. 8 setae near posterior margin and ventral corner per side ( Fig. 10 A). Paramere narrowed basally and apically, slightly curved ventrally, with ca. 7 sensilla distally ( Fig. 10 C). Measurements. BL = 2.47 mm in holotype (range in 2Ƌ paratypes : 2.33–2.40 mm ), THL = 1.20 mm ( 1.10– 1.13 mm ), WL = 1.83 mm ( 1.63–1.70 mm ), WW = 0.97 mm ( 0.87–0.93 mm ), arb = 5/3 (6/2), avd = 0.85 (0.77– 0.83), adf = 2.60 (2.40–2.60), flw = 2.20 (2.20), FW/HW = 0.25 (0.25), ch/o = 0.03 (0.03), prorb = 0.55 (0.56– 0.72), rcorb = 0.60 (0.56–0.72), vb = 0.40 (0.36–0.38), dcl = 0.67 (0.43–0.50), presctl = 0.58 (0.36–0.50), sctl = 1.31 (1.52–1.60), sterno = 0.85 (0.75–0.80), orbito = 1.00 (1.00), dcp = 0.28 (0.29–0.33), sctlp = 0.92 (0.77–0.85), C = 1.55 (1.71–1.75), 4c = 2.11 (1.82–1.90), 4v = 3.16 (2.55–2.80), 5x = 1.46 (1.67–1.80), ac = 2.67 (3.08–3.45), M = 1.00 (0.90–0.91), C3F = 0.73 (0.76–0.78). Type specimens. Holotype Ƌ ( SCAU , No. 128861), CHINA : Wangtianshu , Mengla , Yunnan , 600m , 13.iv.2010 , ex tussock, L Wang . Paratypes : CHINA : 1Ƌ ( SCAU , No. 128862), same data as holotype ; 1Ƌ ( SCAU , No. 128863), Hesong , Menghai , Yunnan , 1750m , 2.iv.2011 , ex moss, SJ Yan. Distribution. China ( Yunnan ). Etymology. A combination of the Latin words “ quadrus ” (= four) + “ cuspidatus ” (= acute), referring to the aedeagus distolaterally with four acute processes.