A mountain of millipedes XI. The trachystreptoform spirostreptids of the Udzungwa Mountains, Tanzania (Diplopoda, Spirostreptida, Spirostreptidae) Author Enghoff, Henrik 9B9D901F-D6C8-4BCA-B11B-CF6EE85B16DC Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, DK- 2100 Copenhagen Ø, Denmark. henghoff@snm.ku.dk Author Ngute, Alain S. K. 4F621C77-C4E7-49C8-ABB2-C6FF1DFD60C6 Forest Research Institute, University of the Sunshine Coast, Sippy Downs, Queensland, Australia. Alain.Ngute@research.usc.edu.au Author Kwezaura, Revocatus L. 2F3825C9-B029-4C61-A0BB-F754D043873C Reforest Africa, Mang’ula, Kilombero District, Tanzania. rauliean@gmail.com Author Laizzer, Richard L. 575788A0-E532-4292-A9D5-F21DAE7742E6 Udzungwa Ecological Monitoring Centre, Udzungwa Mountains National Park, Mang’ula, Kilombero District, Tanzania. richardlowassary@gmail.com Author Lyatuu, Herman M. 7659F933-34F5-4D57-A2E4-F1DBCAC8186E Reforest Africa, Mang’ula, Kilombero District, Tanzania. lyatuuherman@gmail.com Author Mhagawale, Waziri E485EE38-FFDB-4859-8679-CFA20A5FFD3F Reforest Africa, Mang’ula, Kilombero District, Tanzania. mhagawalewaziri@gmail.com Author Mnendendo, Hamidu R. 02489B44-5503-42BF-8DD9-1F40930F0D4D Reforest Africa, Mang’ula, Kilombero District, Tanzania. hamidumnendendo@gmail.com Author Marshall, Andrew R. 0AE8656F-DA17-4955-95A9-1B9EF7910037 Forest Research Institute, University of the Sunshine Coast, Sippy Downs, Queensland, Australia. & Reforest Africa, Mang’ula, Kilombero District, Tanzania. & Flamingo Land Ltd., Kirby Misperton, North Yorkshire, UK. amarsha1@usc.edu.au text European Journal of Taxonomy 2024 2024-01-22 918 1 1 50 https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2405/10603 journal article 10.5852/ejt.2024.918.2405 2118-9773 10554532 8E007E74-FB24-422F-AFA8-033A2D172839 Attemsostreptus cataractae Enghoff sp. nov. urn:lsid:zoobank.org:act: 7D1B998B-D832-4821-88FF-B2F7904B3083 Figs 2 , 8 Diagnosis Differs from congeners, except A. costatus , by the very long (> half length of gonopod coxa), only moderately curved and almost straightly laterad lateroapical metaplical process ( lap ). Differs from A. costatus by a somewhat shorter lap and by the lack of a distinct, rounded mesapical metaplical process ( map ) – compare Fig. 8D with fig. 1 in Akkari & Enghoff (2019) . Etymology A Latin noun in the genitive, meaning ‘of the waterfall’ and referring to the type locality. The name was suggested, but never published, by Richard L. Hoffman (1927–2012), great American myriapodologist. Material examined (total 1 ♂ , 1 ♀ ) Holotype TANZANIA • ♂ ; Morogoro Region , Kilombero District , base of Sanje Falls , Mwanihana Forest Reserve ; Apr. 1984 ; K.M. Howell leg.; VMNH 112021 . Paratype TANZANIA • 1 ♀ ; same collection data as for holotype; VMNH 112022 . Description (male) SIZE . Length unknown (specimen in several pieces, collum and rings 2–6 missing). Diameter 4.2 mm . 54 podous rings (including 1–6), no apodous rings in front of telson. COLOUR . After 39 years in alcohol antennae, legs and head below antennae yellowish; head between antennae brown, marbled in upper part. Collum marbled brown, with brown anterior band expanded in middle. Body rings faded to whitish tallow except for amber-coloured posterior ¼ of metazonites. HEAD . Eyes each with ca 30 ommatidia in ca 12 vertical and 5 horizontal rows, extending just mesad of antennal sockets. Antennae (presumably) reaching back to ring 4 when stretched. COLLUM . With three strong complete carinae, otherwise unornamented, resembles Fig. 9A ; lateral lobes slightly expanded. BODY RINGS ( Fig. 8A–C ). Anterior part of prozonites with dense transverse microsculpture which posteriorly gives way to an at first irregular, then regular cell structure; in dorsal part of ring, surface of last cell row coarsely pitted. Metazonites slightly vaulted, carinate throughout; in dorsal part of ring surface between ridges coarsely pitted. Suture between pro- and metazonites simple, straight. Ozopores ( oz ) between two ridges, in middle of metazonite. No sigilla. TELSON . Similar to that of A. reflexus ( Fig. 4B–D ). Fig. 8. Attemsostreptus cataractae Enghoff sp. nov. , holotype, ♂ (VMNH 112021). A–C . Midbody rings. A . Four rings with legs. B . Close-up of metazonite and posterior part of prozonite. C . Closeup of prozonite and anterior part of metazonite. D–G . Right gonopod. D . Anterior view. E . Anteriormesal view. F . Distal part of telopodite. G . Tip of telopodite. Abbreviations: atp = antetorsal process (highlighted in yellow in D); lap = lateroapical metaplical process; map = mesapical metaplical process; mp = metaplica; mr = metaplical “roof”; oz = ozopore (hard to see); pp = proplica; px = paracoxite; sl = serrate lamella; slm = solenomere. Scale bars: A = 1 mm; B–F = 0.2 mm; G = 0.05 mm. LEGS (leg-pairs 1–7 missing). Length ca 0.9×body diameter. Postfemora and tibiae with ventral pads; pads decreasing in size towards posterior and absent from last pairs of legs. GONOPOD COXA ( Fig. 8D–E ). Slender, parallel-sided. Proplica ( pp ) apically punctate and slightly expanded, apical margin oblique, fitting under metaplical roof ( mr ), mesally ending in acute-angled process. Metaplica ( mp ) without distomesal spine; apically bent over and forming a ‘roof’ ( mr ) over the gonocoel; mesally slightly produced into poorly defined mesapical process ( map ); laterally forming a very long (>half length of entire coxa) slender process ( lap ); process directed laterad, almost straight except for ‘downwards’-bent tip. GONOPOD TELOPODITE ( Fig. 8D–G ). With a very long, slender, pointed antetorsal process ( atp ) in the shape of a large hook. Post-torsal part of telopodite very slender, directed basad, describing a loose 360° spiral, extending past base of coxa where it makes a 180° turn such that the last part is directed apicad; tip divided into lamellar, slightly serrate process ( sl ) and pointed solenomere ( slm ). Descriptive notes (female) Length 75 mm . Diameter 5.5 mm , 54 podous rings, no apodous ring in front of telson. Lateral lobes of collum not expanded. Other non-sexual characters as in male. Distribution and habitat Known only from the type locality; “base of Sanje Falls” could be anywhere between 300 and 500 m a.s.l.