<document id="89ABDABF5A496D1BB418A7FE884433FB" ID-DOI="10.5852/ejt.2024.918.2405" ID-ISSN="2118-9773" ID-ZooBank="8E007E74-FB24-422F-AFA8-033A2D172839" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1705999280996" checkinUser="plazi" docAuthor="Enghoff, Henrik, Ngute, Alain S. K., Kwezaura, Revocatus L., Laizzer, Richard L., Lyatuu, Herman M., Mhagawale, Waziri, Mnendendo, Hamidu R. &amp; Marshall, Andrew R." docDate="2024" docId="714A87FAFFF4E072CF0FFE0BAB90EA45" docLanguage="en" docName="EJT.2024.918.1-50.pdf" docOrigin="European Journal of Taxonomy 918 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2405/10603" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Attemsostreptus Verhoeff 1941" docType="treatment" docVersion="1" lastPageNumber="6" masterDocId="8D73FF82FFF1E077CD3AFFF7AA3DEF08" masterDocTitle="A mountain of millipedes XI. The trachystreptoform spirostreptids of the Udzungwa Mountains, Tanzania (Diplopoda, Spirostreptida, Spirostreptidae)" masterLastPageNumber="50" masterPageNumber="1" pageNumber="6" updateTime="1705999410232" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="626D5331F950F2B46B52663D28F398FC">A mountain of millipedes XI. The trachystreptoform spirostreptids of the Udzungwa Mountains, Tanzania (Diplopoda, Spirostreptida, Spirostreptidae)</mods:title>
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<mods:namePart id="0DC62AD1F6AC8EDB2BF2BC3D30800E01">Enghoff, Henrik</mods:namePart>
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<mods:namePart id="69D7FDB1B9B9D94516035729EBCEE9FD">Ngute, Alain S. K.</mods:namePart>
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<mods:affiliation id="1F88D2EE253BE0746172BC89FE91EA57">Reforest Africa, Mang’ula, Kilombero District, Tanzania.</mods:affiliation>
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<mods:namePart id="F76ECD0B549110D68E1631FB48A7E96E">Laizzer, Richard L.</mods:namePart>
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<mods:affiliation id="13D1FF10502964C77C0F672449FB6C75">Udzungwa Ecological Monitoring Centre, Udzungwa Mountains National Park, Mang’ula, Kilombero District, Tanzania.</mods:affiliation>
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<mods:namePart id="E209731D246E7E2252E6744311D27EC7">Lyatuu, Herman M.</mods:namePart>
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<mods:namePart id="DA9EF8314ED753F2AC90E5E6A9D906FC">Mnendendo, Hamidu R.</mods:namePart>
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<mods:affiliation id="4643B7B6F57321C142A5998BF05959B8">Forest Research Institute, University of the Sunshine Coast, Sippy Downs, Queensland, Australia. &amp; Reforest Africa, Mang’ula, Kilombero District, Tanzania. &amp; Flamingo Land Ltd., Kirby Misperton, North Yorkshire, UK.</mods:affiliation>
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Genus 
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<emphasis id="CB97EAFEFFF4E072CFBDFE0BA97AED1E" bold="true" box="[647,839,508,534]" italics="true" pageId="5" pageNumber="6">Attemsostreptus</emphasis>
<bibRefCitation id="9D724B1DFFF4E072CE75FE0BA9C2ED1E" author="Verhoeff K. W." box="[847,1023,508,534]" pageId="5" pageNumber="6" pagination="231 - 274" refId="ref23255" refString="Verhoeff K. W. 1941. Studien an athiopischen Diplopoden. Jenaische Zeitschrift fur Naturwissenschaft 73: 231 - 274." type="journal article" year="1941">Verhoeff, 1941</bibRefCitation>
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<taxonomicName id="3EE34D6FFFF4E072CD87FDC9A854ED5F" ID-CoL="7NRSX" authority="Verhoeff, 1941: 261" authorityName="Verhoeff" authorityPageNumber="261" authorityYear="1941" box="[189,617,573,600]" class="Diplopoda" family="Spirostreptidae" genus="Attemsostreptus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="CB97EAFEFFF4E072CD87FDC9AB4AED50" box="[189,375,574,600]" italics="true" pageId="5" pageNumber="6">Attemsostreptus</emphasis>
<bibRefCitation id="9D724B1DFFF4E072CC44FDCAA854ED5F" author="Verhoeff K. W." box="[382,617,573,599]" pageId="5" pageNumber="6" pagination="231 - 274" refId="ref23255" refString="Verhoeff K. W. 1941. Studien an athiopischen Diplopoden. Jenaische Zeitschrift fur Naturwissenschaft 73: 231 - 274." type="journal article" year="1941">Verhoeff, 1941: 261</bibRefCitation>
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<emphasis id="CB97EAFEFFF4E072CD87FD88AB65ED90" bold="true" box="[189,344,638,665]" pageId="5" pageNumber="6">
<typeStatus id="2658884EFFF4E072CD87FD88AAC7ED91" box="[189,250,639,665]" pageId="5" pageNumber="6">Type</typeStatus>
species
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<taxonomicName id="3EE34D6FFFF4E072CD87FD59A8A8EDCF" ID-CoL="JMX2" authority="Verhoeff, 1941" authorityName="Verhoeff" authorityYear="1941" box="[189,661,685,712]" class="Diplopoda" family="Spirostreptidae" genus="Attemsostreptus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="costatus">
<emphasis id="CB97EAFEFFF4E072CD87FD59ABE2EDCF" box="[189,479,686,712]" italics="true" pageId="5" pageNumber="6">Attemsostreptus costatus</emphasis>
<bibRefCitation id="9D724B1DFFF4E072CCDCFD5AA8A8EDCF" author="Verhoeff K. W." box="[486,661,685,711]" pageId="5" pageNumber="6" pagination="231 - 274" refId="ref23255" refString="Verhoeff K. W. 1941. Studien an athiopischen Diplopoden. Jenaische Zeitschrift fur Naturwissenschaft 73: 231 - 274." type="journal article" year="1941">Verhoeff, 1941</bibRefCitation>
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(
<collectingCountry id="81F4767CFFF4E072CF9EFD5AA92CEDCF" box="[676,785,685,711]" name="Tanzania" pageId="5" pageNumber="6">Tanzania</collectingCountry>
) by monotypy.
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<emphasis id="CB97EAFEFFF4E072CD87FD19ABE5EC00" bold="true" box="[189,472,750,776]" pageId="5" pageNumber="6">Other included species</emphasis>
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<taxonomicName id="3EE34D6FFFF4E072CD87FCE9A8C2EC3F" ID-CoL="846XH" authority="Akkari &amp; Enghoff, 2019" authorityName="Akkari &amp; Enghoff" authorityYear="2019" box="[189,767,797,824]" class="Diplopoda" family="Spirostreptidae" genus="Attemsostreptus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="reflexus">
<emphasis id="CB97EAFEFFF4E072CD87FCE9ABE4EC3F" box="[189,473,797,824]" italics="true" pageId="5" pageNumber="6">Attemsostreptus reflexus</emphasis>
<bibRefCitation id="9D724B1DFFF4E072CCE4FCEAA8C2EC3F" author="Akkari N. &amp; Enghoff H." box="[478,767,797,823]" pageId="5" pageNumber="6" pagination="1 - 12" refId="ref20811" refString="Akkari N. &amp; Enghoff H. 2019. Revision of the genus Attemsostreptus Verhoeff, 1941 with description of a new species from Tanzania and notes on the tribe Trachystreptini. European Journal of Taxonomy 575: 1 - 12. https: // doi. org / 10.5852 / ejt. 2019.575" type="journal article" year="2019">Akkari &amp; Enghoff, 2019</bibRefCitation>
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<taxonomicName id="3EE34D6FFFF4E072CD87FCB6A85EEC52" authority="Enghoff" authorityName="Enghoff &amp; Ngute &amp; Kwezaura &amp; Laizzer &amp; Lyatuu &amp; Mhagawale &amp; Mnendendo &amp; Marshall" authorityYear="2024" box="[189,611,832,859]" class="Diplopoda" family="Spirostreptidae" genus="Attemsostreptus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="cataractae" status="sp. nov.">
<emphasis id="CB97EAFEFFF4E072CD87FCB6ABC1EC52" box="[189,508,833,859]" italics="true" pageId="5" pageNumber="6">Attemsostreptus cataractae</emphasis>
Enghoff
</taxonomicName>
<taxonomicNameLabel id="D0A45785FFF4E072CF50FCB6A8F8EC53" box="[618,709,833,859]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
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<taxonomicName id="3EE34D6FFFF4E072CD87FC93A851EC75" authority="Enghoff" authorityName="Enghoff &amp; Ngute &amp; Kwezaura &amp; Laizzer &amp; Lyatuu &amp; Mhagawale &amp; Mnendendo &amp; Marshall" authorityYear="2024" box="[189,620,867,894]" class="Diplopoda" family="Spirostreptidae" genus="Attemsostreptus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="julostriatus" status="sp. nov.">
<emphasis id="CB97EAFEFFF4E072CD87FC93A838EC75" box="[189,517,867,894]" italics="true" pageId="5" pageNumber="6">Attemsostreptus julostriatus</emphasis>
Enghoff
</taxonomicName>
<taxonomicNameLabel id="D0A45785FFF4E072CF49FC93A8F0EC76" box="[627,717,868,894]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="F95C36ECFFF4E072CD87FC70A8FEECA9" blockId="5.[189,767,797,929]" box="[189,707,903,929]" pageId="5" pageNumber="6">
<taxonomicName id="3EE34D6FFFF4E072CD87FC70A85FECA9" authority="Enghoff" authorityName="Enghoff &amp; Ngute &amp; Kwezaura &amp; Laizzer &amp; Lyatuu &amp; Mhagawale &amp; Mnendendo &amp; Marshall" authorityYear="2024" box="[189,610,903,929]" class="Diplopoda" family="Spirostreptidae" genus="Attemsostreptus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="leptoptilos" status="sp. nov.">
<emphasis id="CB97EAFEFFF4E072CD87FC70ABC6ECA9" box="[189,507,903,929]" italics="true" pageId="5" pageNumber="6">Attemsostreptus leptoptilos</emphasis>
Enghoff
</taxonomicName>
<taxonomicNameLabel id="D0A45785FFF4E072CF53FC70A8FEECA9" box="[617,707,903,929]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="B1F96567FFF4E072CD87FC3FAB90EA45" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="F95C36ECFFF4E072CD87FC3FAB0BECEA" blockId="5.[189,310,968,994]" box="[189,310,968,994]" pageId="5" pageNumber="6">
<heading id="A2148180FFF4E072CD87FC3FAB0BECEA" bold="true" box="[189,310,968,994]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="CB97EAFEFFF4E072CD87FC3FAB0BECEA" bold="true" box="[189,310,968,994]" pageId="5" pageNumber="6">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="F95C36ECFFF4E072CD87FC00AB90EA45" blockId="5.[188,1400,1015,1357]" pageId="5" pageNumber="6">
The discovery of three new species in the genus necessitates a revision of the genus diagnosis proposed by 
<bibRefCitation id="9D724B1DFFF4E072CDDCFBEDA81AEB3C" author="Akkari N. &amp; Enghoff H." box="[230,551,1050,1076]" pageId="5" pageNumber="6" pagination="1 - 12" refId="ref20811" refString="Akkari N. &amp; Enghoff H. 2019. Revision of the genus Attemsostreptus Verhoeff, 1941 with description of a new species from Tanzania and notes on the tribe Trachystreptini. European Journal of Taxonomy 575: 1 - 12. https: // doi. org / 10.5852 / ejt. 2019.575" type="journal article" year="2019">Akkari &amp; Enghoff (2019)</bibRefCitation>
. Species of 
<taxonomicName id="3EE34D6FFFF4E072CFFEFBEDA943EB3C" authorityName="Verhoeff" authorityYear="1941" box="[708,894,1050,1076]" class="Diplopoda" family="Spirostreptidae" genus="Attemsostreptus" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="CB97EAFEFFF4E072CFFEFBEDA943EB3C" box="[708,894,1050,1076]" italics="true" pageId="5" pageNumber="6">Attemsostreptus</emphasis>
</taxonomicName>
differ from other 
<taxonomicName id="3EE34D6FFFF4E072C953FBEDAF26EB3C" authorityName="Brandt" authorityYear="1833" box="[1129,1307,1050,1076]" class="Diplopoda" family="Spirostreptidae" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="family">Spirostreptidae</taxonomicName>
by the combination of the following characters: anterior margin of collum unmodified, metazonites with longitudinal ridges (carinae), or at least striae along the entire perimeter of the body (trachystreptoform habitus); prefemoral lobes of first pair of male legs triangular, covered with minute short spines; tip of gonopod coxal metaplica folded back on anterior side, forming a ‘roof’ or canopy over gonocoel; a long to very long pointed lateroapical metaplical process; gonopod telopodite long and slender, with a long to very long, pointed antetorsal process; posttorsal part of telopodite very long and thin, forming loose coils; telopodite ending in two short branches of subequal length, one broad and more or less serrated, the other acuminate.
</paragraph>
</subSubSection>
</treatment>
</document>