The mostly cavernicolous millipede genus Stygiiulus Verhoeff, 1929, stat. nov.: taxonomy, distribution and phylogenetic relationships (Diplopoda, Julida, Julidae) Author Vagalinski, Boyan CF6A5691-5C32-4CFF-9EB6-045DEB16F289 Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, 2 Gagarin Street, 1113, Sofia, Bulgaria. boyanv84@gmail.com Author Borissov, Simeon C035E1AC-D4FB-4A3F-8B38-5D38A9665E88 Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, 2 Gagarin Street, 1113, Sofia, Bulgaria. borissovsb@gmail.com Author Bobeva, Aneliya 0F4EA315-46BE-49AC-8FB4-CF710C5B2451 Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, 2 Gagarin Street, 1113, Sofia, Bulgaria. aneliabobeva@gmail.com Author Canciani, Giacomo 4BB21FD2-15C8-44BD-907A-4498D7D02A15 Jamarski Klub Kraški Krti / G. S. Talpe del Carso, loc. San Michele del Carso / Vrh, I- 34070 Savogna d’Isonzo (GO), Italy. & Associazione Speleologica Forum Julii Speleo, Via Carraria 99, I- 33043 Cividale del Friuli (UD), Italy. giacomo.canciani90@gmail.com Author Antić, Dragan Ž. 2368C980-2042-4137-998D-8C91E4337185 University of Belgrade - Faculty of Biology, Institute of Zoology, Studentski Trg 16, 11000 Belgrade, Serbia. & Serbian Biospeleological Society, Trg Dositeja Obradovića 2, 21000 Novi Sad, Serbia. dragan.antic@bio.bg.ac.rs text European Journal of Taxonomy 2022 2022-02-28 798 30 69 http://dx.doi.org/10.5852/ejt.2022.798.1669 journal article 20422 10.5852/ejt.2022.798.1669 31fec661-5dac-4626-856d-4ba4680efee7 2118-9773 6323002 50692D26-A41C-4F85-B207-A6747FD07470 Stygiiulus maximus ( Verhoeff, 1929 ) comb. nov. Fig. 10D Mesoporoiulus maximu s Verhoeff, 1929: 19–20 , fig. 2. Typhloiulus ( Stygiiulus ) maximus – Verhoeff 1930: 9 , 12–13, figs 1–2. — Manfredi 1932: 81 . Typhloiulus ( Mesoporoiulus ) maximus – Pretner & Strasser 1931: 87 . Typhloiulus maximus var. longicauda Strasser, 1962: 59–60 , fig. 71. Typhloiulus maximus var. maximus – Strasser 1962: 59 , fig. 72. Typhloiulus maximus – Attems 1949: 145 . — Strasser 1971a: 14 . — Vagalinski et al. 2015: 342–343 . not Typhloiulus tobias – Attems 1927: 250–251 , figs 352–354. Diagnosis A species of Stygiiulus stat. nov. with normal mouthparts. Distinguishable from congeners by the combination of certain gonopodal characters ( Fig. 10D ), viz., a mostly straight pro- and mesomere, and an opisthomere with a faint and blunt posterior hump, a marginally broad (not tapering) and deeply serrated velum, and a solenomere with both the anterior and the posterior solenomeral branch being well developed and clearly discernible. Distribution This species has the widest distribution of all representatives of the genus Stygiiulus stat. nov. Known from numerous caves, as well as epigean habitats, ranging from the Julian Alps in the east, across the southern parts of the Carnic Alps Range, through the Venetian Prealps, all the way to the Piave River in the west ( Fig. 13 , pink squares). Remark Attems (1927: 250 , 251, figs 352–354) gave a short description and drawings of what he thought was already described as S. tobias comb. nov. from Monte Cavallo ( Lombardy ). Just two/three years later, Verhoeff (1930) described another blind julid, S. maximus comb. nov. , from a cave in the same area. What is evident from the gonopod drawings of both Attems and Verhoeff alone – that is that Attems’ (1927) record actually refers to S. maximus comb. nov. – was already confirmed by Strasser (1962: 60) based on re-examination of the specimens from Monte Cavallo.