New data on the wolf spiders from theAcantholycosa-complex (Araneae: Lycosidae) from the South Siberia
Author
Fomichev, Alexander A.
Altai State University, Lenina Pr., 61, Barnaul, RF- 656049, Russia. & Tomsk State University, Lenina Pr., 36, Tomsk, RF- 634050, Russia.
text
Zootaxa
2021
2021-08-27
5026
4
567
585
journal article
10.11646/zootaxa.5026.4.7
1175-5326
5300792
E712BC8E-3E75-422C-8587-69ED17B6CDD5
Acantholycosa levinae
Marusik, Azarkina & Koponen, 2004
Figs 4–5
,
18–19
, 28–31, 36–37, 66–68, 75–76
A. levinae
Marusik, Azarkina & Koponen, 2004: 112
, f. 61–65 (
♂
).
A. levinae
—
Marusik & Omelko, 2017: 597
, f. 11 (
♂
).
Material examined.
RUSSIA
:
Altai Republic
:
1♂
1♀
(
ISEA
, 001.8658)
,
1♀
(
ISEA
, 001.8722),
Katunskyi Mt. Range
,
Srednee Mul’tinskoe Lake
,
49°59’N
,
85°50’E
, scree and lichen pillows,
1650–1760 m
, 23–
27.07.2019
,
A.A. Fomichev
,
Yu.V. Dyachkov
;
2♀
(
ISEA
, 001.8659),
Katunskyi Mt. Range
, watershed of
Mul’ta River
and
Kuiguk River
,
49°59.019’N
,
85°51.096’E
, scree,
2400 m
,
26.07.2019
,
A.A. Fomichev
,
Yu.V. Dyachkov, E.A
.
Nepaeva
;
3♂
1♀
(
ZMMU
)
,
Katunskyi Mt. Range
,
Poperechnoe Lake
,
49°55’N
,
85°53’E
, scree,
1885–2000 m
,
21.07.2018
,
A.E. Naidenov
,
K.E. Naidenova
.
Diagnosis.
Acantholycosa levinae
resembles
A. baltoroi
. Both species have protruding palea (
Pl
) with triangleshaped projection (
Ts
), robust terminal apophysis (
Ta
) and relatively straight embolus (
Em
). The males of these two species differ by the tip of the embolus (bilobated in
A. levinae
, and single-blade in
A. baltoroi
) and by the shape of the terminal apophysis (curved and bifurcated in
A. levinae
, and straight and blunt in
A. baltoroi
, cf. Fig. 29 and fig.
60 in
Marusik
et al.
2004
). The female of
A. levinae
differs from those of
A. baltoroi
by the shape of epigynal fovea (
Fo
), which is rectangular in the former species (vs. heart-shaped) and by rectangular septal base (
Sb
) (vs. anchor-shaped, cf.
Fig. 66
and fig. 2D in
Buchar 1976
). The epigyne of
A. levinae
is similar to those of species from the
plumalis
-group:
A. logunovi
Marusik, Azarkina & Koponen, 2004
,
A. paraplumalis
Marusik, Azarkina & Koponen, 2004
and
A. plumalis
Marusik, Azarkina & Koponen, 2004
. The similarity includes a rectangular shape of the fovea as well as septal base and apical pocket (
Ap
) consisting of one hood. The female of
A. levinae
differs from all aforementioned species by a reduced septal stem. In addition, it can be separated from
A. logunovi
and
A. paraplumalis
by having the apical pocket as wide as the septal base (vs. twice narrower). Finally, the female of
A. levinae
can be distinguished from
A. plumalis
by having receptacles (
Re
) almost touching each other (vs. widely spaced, cf.
Fig. 66, 68
and fig.
3 in
Fomichev & Marusik 2011
, and figs 47, 50–
52 in
Marusik
et al.
2004
).
Description.
Female.Total length 8.6. Carapace: 4.15 long, 3.3 wide. General appearance as in
Fig. 5
.Coloration. Carapace, sternum and labium dark brown. Eye field black. Chelicerae, endites and palps yellow-brown. Coxae yellow-gray. Legs yellow-brown, femora with transverse dark gray stripes. Abdomen dark gray dorsally, with dark brown cardiac mark. Venter of abdomen yellow-gray. Spinnerets dark gray. Spination of leg I: Fe d1-1-1 p0-0-2 r0- 1-1; Ti p1-0-0 r1-0-0 v1-2-2-2-2-2; Mt p0-1-0 r0-1-0 v2-2-0. Leg measurements: I 3.9, 1.8, 3.65, 3.45, 1.7 (14.5); II 3.9, 1.75, 3.4, 3.55, 1.75 (14.35); III 3.8, 1.6, 3.15, 4.3, 1.85 (14.7); IV 4.85, 1.8, 4.3, 6.85, 2.55 (20.35).
Epigyne and internal genitalia as shown in
Figs 66–68
. Fovea as wide as long. Septal base smoothly declines anteriorly. Lips of the epigyne (
Li
) separated by one of their wide. Receptacles clavate, not clearly delineated from copulatory ducts (
Cd
). Copulatory ducts 1.5 times longer than receptacles.
Male. See
Marusik
et al.
(2004)
for description. General appearance as in
Fig. 4
. Palp as in
Figs 18–19
, 28–31, 36–37. Tegular apophysis (
Tg
) almost oval, with reduced apical and basal arms. Palea with abrupt retrolateral margin. Triangle shaped projection almost straight, with a hooked apical part. Terminal apophysis massive, subdivided into two lobes. Embolus with a spine (
Sp
) in the basal part. Distal part of the embolus subdivided into triangular secondary part and embolus proper.
Distribution.
Known from several close localities in the Katunskyi Mt. Range in
Altai
Mountains (
Figs 75– 76
).
Comments.
There are some doubts if the females match with males due to the morphology of the epigyne. Since
A. levinae
belongs to the
baltoroi
-group, this species epigyne would be expected to have an anchor-shaped septal base. Instead, the epigyne of the females described herein are similar to those of species from the
plumalis
-group. Nevertheless, males and females were collected together in two different places, thus I treat them as conspecific.