The American Goblin Spiders Of The New Genus Escaphiella (Araneae, Oonopidae)
Author
Platnick, Norman I.
Author
Dupérré, Nadine
text
Bulletin of the American Museum of Natural History
2009
2009-09-03
2009
328
1
151
http://www.bioone.org/doi/abs/10.1206/679.1
journal article
10.1206/679.1
0003-0090
12776972
Escaphiella tonila
,
new species
Figures 269–290
; map 5
TYPES
:
Male
holotype
and female
allotype
taken
2 miles
S of Tonila
,
Colima
,
Mexico
(
Aug. 28, 1965
;
W. Gertsch
,
R. Hastings
), deposited in
AMNH
(PBI_OON 36863)
.
ETYMOLOGY
: The specific name is a noun in apposition taken from the
type
locality.
DIAGNOSIS
: Whether males of this species show palpal asymmetry is unknown; the only available specimen is missing the left palp, but has a distinctively rebordered posterior margin on the endite excavation (figs. 270, 276), a club-shaped palpal femur (fig. 277), and an extremely long, thin embolus (fig. 280). Females have a distinctively widened anterior epigynal sclerite containing an S-shaped darkening (fig. 290), and both sexes seem to have slightly reduced eyes (figs. 272, 288).
MALE
(PBI_OON 36863): Total length 1.17 (figs. 269, 270, 273). Carapace pale orange, surface of elevated portion of pars cephalica smooth, sides granulate; lateral margins without denticles (figs. 271, 274). Eyes: ALE circular, PME squared; ALE separated by less than their radius, ALE-PLE touching, PME touching throughout most of their length, PLE-PME touching (fig. 272). Sternum pale orange, without radial furrows between coxae I–II, II–III, III–IV, surface smooth, microsculpture absent (fig. 276). Chelicerae, endites, and labium pale orange. Posterior margin of endite excavation rebordered (fig. 276). Abdomen ovoid; book lung covers large, ovoid; pedicel tube medium (fig. 275), scutum extending far dorsal of pedicel; dorsal scutum pale orange, covering full length of abdomen, no soft tissue visible from above, middle surface smooth, sides smooth; postepigastric scutum pale orange, sides smooth. PMS not observed. Legs pale orange. Palp with proximal segments pale orange; embolus dark, greatly elongated (figs. 278–281); femur enlarged, club shaped (fig. 277); cymbium and bulb pale orange, subdistal row of six long, stiff setae marking edge of cymbial area on prolateral side (fig. 279).
Figs. 240–248.
Escaphiella catemaco
,
new species
, male.
240.
Habitus, dorsal view.
241.
Same, ventral view.
242.
Carapace, dorsal view.
243.
Habitus, anterior view.
244.
Same, lateral view.
245.
Carapace, lateral view.
246.
Endite, lateral view.
247.
Cephalothorax, ventral view.
248.
Epigastric region, ventral view.
Figs. 249–257.
Escaphiella catemaco
,
new species
, male.
249.
Left palp, dorsal view.
250.
Same, prolateral view.
251.
Same, ventral view.
252.
Same, retrolateral view.
253.
Right palp, dorsal view.
254.
Same, prolateral view.
255.
Same, ventral view.
256.
Same, retrolateral view.
257.
Abdomen, ventral view.
Figs. 258–268.
Escaphiella catemaco
,
new species
, female.
258.
Habitus, dorsal view.
259.
Same, ventral view.
260.
Carapace, dorsal view.
261.
Habitus, anterior view.
262.
Same, lateral view.
263.
Abdomen, ventral view.
264.
Same, anterior view.
265.
Anterior portion of abdomen, lateral view.
266.
Cephalothorax, ventral view.
267.
Epigynum, ventral view.
268.
Carapace, lateral view.
Figs. 269–275.
Escaphiella tonila
,
new species
, male.
269.
Habitus, dorsal view.
270.
Same, ventral view.
271.
Carapace, dorsal view.
272.
Habitus, anterior view.
273.
Same, lateral view.
274.
Carapace, lateral view.
275.
Epigastric region, ventral view.
Figs. 276–281.
Escaphiella tonila
,
new species
, male.
276.
Cephalothorax, ventral view.
277.
Right palpal femur, retrolateral view.
278.
Right palp, dorsal view.
279.
Same, prolateral view.
280.
Same, ventral view.
281.
Same, retrolateral view.
FEMALE
(PBI_OON 36863): Total length 1.28 (figs. 284–286, 289). Abdomen with soft portions of dorsum white (figs. 282, 288); pedicel tube ribbed, scutopedicel region unmodified. PMS present as single median projection (light microscope). Anterior epigynal sclerite containing S-shaped darkening (figs. 283, 287, 290).
Figs. 282–290.
Escaphiella tonila
,
new species
, female.
282.
Habitus, dorsal view.
283.
Same, ventral view.
284.
Cephalothorax, ventral view.
285.
Habitus, lateral view.
286.
Carapace, lateral view.
287.
Abdomen, ventral view.
288.
Habitus, anterior view.
289.
Carapace, dorsal view.
290.
Epigynum, ventral view.
Map 5. Records of
Escaphiella tonila
(star) and
E. chiapa
(circles).
OTHER
MATERIAL
EXAMINED
:
One female
taken with the
types
(AMNH PBI_OON 36863).
DISTRIBUTION
: Known only from the
type
locality in
Colima
,
Mexico
(map 5).