A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae) Author Enghoff, Henrik Zoological Museum, Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, DK- 2100, Copenhagen Ø, Denmark & Corresponding author: henghoff @ snm. ku. dk & urn: lsid: zoobank. org: author: FB 09 A 817 - 000 D- 43 C 3 - BCC 4 - 2 BC 1 E 5373635 henghoff@snm.ku.dk Author Frederiksen, Sara B. 6DE62DD5-0055-424A-A136-0EF290AF5AFD Zoological Museum, Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, DK- 2100, Copenhagen Ø, Denmark & Department of Ecology, Animal Ecology, University of Marburg, Karl-von-Frisch-Strasse 8, D- 35032 Marburg, Germany & Email: sara. frederiksen @ snm. ku. dk & urn: lsid: zoobank. org: author: 6 DE 62 DD 5 - 0055 - 424 A-A 136 - 0 EF 290 AF 5 AFD sara.frederiksen@snm.ku.dk text European Journal of Taxonomy 2015 2015-11-06 150 1 25 journal article 22375 10.5852/ejt.2015.150 f492ad9b-a83d-4d8e-bdf9-20ef759b75c1 2118-9773 3787588 D546A676-35F8-4025-8660-F123A0BA67F7 Aquattuor udzungwensis Enghoff sp. nov. urn:lsid:zoobank.org:act: 60F38BFF-E5EA-4BB0-A691-263DB097502B Figs 3 , 8 Diagnosis Diameter 1.44–1.69 mm , 44–50 podous rings. Gonopods: lateral incision of coxa ( li ) deep with rounded bottom, curving distad; apical palette ( pa ) rounded-triangular, not longer than broad; telomere describing a c. 270° curve in almost one plane; mesal-posterior lamella of telomere ( mpl ) high, basally set off by 45–90° angle; anterior margin of telomere without a triangular tooth; telomere tip with smooth to microspiculate margins, internal surface microspiculate. Etymology The name is a Latin adjective derived from the Udzungwa Mts, home of this (and several other) Aquattuor species. Material studied (total: 21 ♂♂ ) Holotype TANZANIA : ♂ , Udzungwa Mts , Iringa Region , Udzungwa Scarp FR , above Chita village , 750 m asl , 23 Oct. –14 Nov. 1984 , lowland rain forest, N. Scharff leg. ( ZMUC ). Fig. 8. Aquattuor udzungwensis Enghoff sp. nov. A–B, D. Paratype. C. Specimen from West Kilombero Scarp FR. A . Gonopods in situ . B . Telopodite, ventral view. C . Right gonopod, anterior view. D . Telomere tip. li = lateral incision; pa = palette; mpl = mesal-posterior lamella of telomere. Scale bars: A–C = 0.1 mm; D = 0.05 mm. Paratypes TANZANIA : 4 ♂♂ , same data as holotype , but 25–29 Oct. 1984 , pitfall traps (ZMUC). Referred non-type material TANZANIA : Iringa Region , Iringa District , Udzungwa Mts, West Kilombero Scarp FR, montane forest, Frontier Tanzania coll.: 1 ♂ , 07°53’19.5” S , 36°23’11.6” E , 1100–1300 m asl, Plot Ukami, 30 Nov. 2000 (ZMUC); 6 ♂♂ , 07°53’19.5” S , 36°23’11.6” E , 1111 m asl, 23 Nov. 2000 (ZMUC); 7 ♂♂ , 07°53’19.5” S , 36°23’11.6” E , 1126 m asl, plot 17 – Ukami, 25 Nov. 2000 (ZMUC); 1 ♂ , 07°50’38.4” S , 36°22’17.6” E , 1390 m asl, 13 Nov. 2000 (ZMUC); 1 ♂ , 07°50’38.4” S , 36°22’17.6” E , 1390–1410 m asl, Plot Paradiso, 18 Nov. 2000 (ZMUC). Type locality TANZANIA : Udzungwa Mts, Iringa Region , Udzungwa Scarp FR, above Chita village, 750 m asl, lowland rain forest. Colouration After 15–31 years in alcohol faded, pale yellowish, head and especially collum contrastingly darker. Traces of paramedian, dark bands flanking a middorsal, pale stripe. Distribution and habitat Known from Udzungwa Scarp FR and from West Kilombero Scarp FR. Altitudinal range: 750–1410 m . Habitat: lowland rain forest and montane rain forest. Coexisting species In Udzungwa Scarp FR, Aquattuor stereosathe Enghoff sp. nov. , Chaleponcus circumvallatus Enghoff, 2014 , C. hamerae Enghoff, 2014 , and C. nikolajscharffi Enghoff, 2014 have been found. In West Kilombero Scarp FR, Aquattuor longipala Enghoff sp. nov. , Chaleponcus netus Enghoff, 2014 , and C. circumvallatus Enghoff, 2014 were found in the same sample as A. udzungwensis Enghoff sp. nov. In addition, C. basiliscus Enghoff, 2014 , C. gracilior Enghoff, 2014 , C. ibis Enghoff, 2014 , and C. tintin Enghoff, 2014 were found in West Kilombero Scarp FR.