The earthworm genus Polypheretima Michaelsen, 1934 (Annelida: Clitellata: Megascolecidae) from Sulawesi, Indonesia, with descriptions of four new species
Author
Fahri
Author
Amaliah, Rizki
Author
Annawaty
Author
Nguyen, Anh D.
text
Raffles Bulletin of Zoology
2017
2017-10-30
65
559
573
journal article
10.5281/zenodo.5357964
2345-7600
5357964
9A17DED6-26FE-44F4-A414-E3C650B16D94
Polypheretima elongata
(
Perrier, 1872
)
(
Fig. 7
)
Perichaeta elongata
Perrier, 1872: 124
Megascolex elongata
Vaillant, 1889: 81
Fig. 7.
Polypheretima elongata
(Michaelsen, 1872)
. A, Spermathecal pores; B, Spermathecae (amp=ampulla, dv=diverticulum); C, Male pore region, ventral view (mp=opening of copulatory pouchs, gm=genital marking); D, Male pore region, lateral view; E, Prostate gland. Scale bar = 1 mm.
Perichaeta biserialis
Perrier, 1875: 1044
Perichaeta acystis
Beddard, 1895: 423
Amyntas elongatus
Beddard, 1900: 650
Pheretima elongata
Michaelsen, 1900: 265
Metapheretima elongata
Sims & Easton, 1972: 233
;
Easton, 1976: 40
(For more synonyms, see
Easton, 1976
)
Material examined.
4 clitellates (
MZB
Oli. 0061), the yard,
Margapura village
(
0°31′2.35″N
,
120°57′45.15″E
), elevation of
5 m
asl
.,
Bolano Lambunu
sub-district,
Parigi Mautong district
,
Central Sulawesi province
,
Indonesia
,
20–21 January 2017
, leg.
Fahri
and
Fitrallisan.
6 clitellates (
UNTAD
Oli. 0007), same data as for the sample MZB
Oli.
0061
.
Description.
External characters
. Body generally cylindrical. Prostomium prolobous. First dorsal pore in 12/13. Setae
68–74 in
v,
70–86 in
vii,
72–88 in
viii,
80–92 in
xxv, and 8–12 between male porophores in xviii. Clitellum annular, xiv–xvi, smooth without setae and dorsal pores. Female pore single, mid-ventral in xiv.
Spermathecal pores small, lateroventrally two paired in 5/6/7. Male porophores highly elevated, large; male pores located inside copulatory pouches in xviii. Genital markings simple, large, presetal, paired in xix–xxi, rarely xxii.
Internal characters
. Septa 3/4/5/6/7/8 thickened, 8/9/10 absent, 10/11/12/13 thin. Gizzard large, round after viii. Last heart in xiii. Intestinal origin at xv; caeca absent. Pharyngeal micronephridia well-developed on septum 7/8. Typhlosole simple.
Spermathecae large, about 1–5 altogether, 1–2 per segment in 5/6 and 1–2 per segment in 6/7 (
Table 6
). The number of spermathecae variable in specimens (
Table 6
). Ampulla large and round; duct relatively short, about 1/6 as long as ampulla. Diverticulum simple, much shorter than ampulla, slightly expanded distad, and attached to the base of duct. No accessory glands.
Holandric. Testes sacs paired in x and xi, small, separated, ventral, yellowish. Seminal vesicles in xi–xii. Ovaries paired in xiii, ovisacs in xiii. Prostate glands paired from xvi–xix, racemose. Prostatic ducts C-shaped, bigger distally, ending at xviii. No accessory glands.
Habitat and ecology.
Specimens were found in the yard in Margapura village, Bolano Lambunu district at
5 m
asl. It was collected from sandy soils near water drainages at depth of
5–20 cm
.
Remarks
.
Polypheretima elongata
is an exotic species widely distributed in
Sulawesi
and other regions in
Indonesia
(
Easton, 1976
). It is also commonly found in the Indochinese region (
Thai, 1983
;
Nguyen et al., 2016
).