Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part XXXI. Two new genera from Somaliland: gen n gen n (Buthidae)
Author
Kovařík, František
text
Euscorpius
2024
386
1
11
journal article
10.5281/zenodo.11449725
1536-9307
11449725
D792C9D7-1879-4006-A5BF-00FFA934F83F
Sanaag
gen
.
n
.
(
Fig. 1
,
Table 1
)
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70E9D5C9-
38FB-4FA8-93C9-FE3709912D94
TYPE
SPECIES
.
Gint maidensis
KovařÍk, Lowe, Just, Awale, Elmi & ŠťÁhlavský, 2018
ETYMOLOGY. The generic name
Sanaag
(masculine) is given after the region of occurrence.
DIAGNOSIS. Total length up to
31–37 mm
(male) or
39–48 mm
(female); carapace trapezoidal, in lateral view preocular area not distinctlY inclined towards anterior margin, level with or higher than postocular area; surface of carapace denselY granular, with onlY anterior median carinae developed; ventral aspect of cheliceral fixed finger with two denticles; tergites denselY granular, with three carinae of which lateral pair on I and II are inconspicuous; sternites III–VI with finelY microdenticulate posterior margins, lacking larger non-contiguous denticles; pectinal tooth number 25–31; pectines with fulcra, hirsute; hemispermatophore with large, tall, subtriangular hook–like basal lobe; metasomal segment I with 8 carinae, ventromedial pair being obsolete; metasoma II–III with 10 carinae; metasoma I ventrallY smooth; metasoma V with enlarged ‘lobate’ dentition on ventrolateral carinae which maY be reduced; telson rather bulbous, more so in males, vesicle with moderate posterior slope, not sharplY inclined or truncated, lacking subaculear tubercle, aculeus shorter or the same as vesicle; all segments of metasoma and pedipalps sparselY hirsute, with long setae in both sexes, dentate margin of movable finger of pedipalp with 9–10 rows of granules, each with one external and one internal accessorY granule, 5 terminal granules (4 terminal and one proximal terminal); trichobothrial pattern orthobothriotaxic
tYpe
A; dorsal trichobothria of femur arranged in b- configuration; pedipalp patella with 7 external trichobothria; patella trichobothrium
d
3
internal to dorsomedian carina; tibial spurs present on legs III–IV.
Figure 1
.
Sanaag maidensis
(KovařÍk et al., 2018)
comb
.
n
.
, male paratYpe in vivo habitus.
|
Sanaag maidensis
|
Sanaag maidensis
|
Sahil elmii
gen
. et
sp
.
n
.
|
|
Dimensions (mm)
|
♂ holotype |
♀ paratype |
♂ holotype |
| Carapace |
L / W |
3.80 / 4.02 |
4.62 / 5.95 |
3.36 / 3.59 |
| Mesosoma |
L |
9.60 |
14.10 |
7.65 |
| Tergite VII |
L / W |
2.90 / 3.95 |
3.75 / 5.75 |
1.80 / 3.54 |
| Metasoma + telson |
L |
23.02 |
29.47 |
17.37 |
| Segment I |
L / W / D |
3.10 / 2.55 / 2.21 |
3.75 / 3.07 / 2.52 |
2.23 / 2.02 / 1.74 |
| Segment II |
L / W / D |
3.55 / 2.42 / 2.17 |
4.35 / 2.85 / 2.60 |
2.63 / 1.84 / 1.81 |
| Segment III |
L / W / D |
3.70 / 2.40 / 2.25 |
4.65 / 2.82 / 2.70 |
2.83 / 1.82 / 1.88 |
| Segment IV |
L / W / D |
4.10 / 2.30 / 2.15 |
5.20 / 2.85 / 2.65 |
3.18 / 1.82 / 1.60 |
| Segment V |
L / W / D |
4.55 / 2.30 / 1.95 |
5.87 / 2.92 / 2.55 |
3.51 / 1.87 / 1.55 |
| Telson |
L / W / D |
4.02 / 1.52 / 1.45 |
5.65 / 2.35 / 2.05 |
2.99 / 1.38 / 1.10 |
| Pedipalp |
L |
12.60 |
16.03 |
8.83 |
| Femur |
L / W |
3.25 / 1.00 |
4.13 / 1.15 |
2.34 / 0.82 |
| Patella |
L / W |
3.95 / 1.35 |
5.05 / 1.75 |
2.57 / 1.02 |
| Chela |
L |
5.40 |
6.85 |
3.92 |
| Manus |
W / D |
1.08 / 1.10 |
1.55 / 1.55 |
0.89 / 0.95 |
| Movable finger |
L |
3.35 |
4.80 |
2.59 |
| Total |
L |
36.43 |
48.20 |
28.38 |
Table 1
. Comparative measurements of tYpes of
Sanaag maidensis
(KovařÍk et al., 2018)
comb
.
n
.
and
Sahil elmii
gen
. et
sp
.
n
.
Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D).
AFFINITIES.
Sanaag
gen
.
n
.
belongs to the ‘
Buthus
’ group of
Fet et al., 2005
(
ŠtundlovÁ et al., 2022
), according to the following characters: trichobothrial pattern
tYpe
A-β; patella trichobothrium
d
3 internal to dorsomedian carina, manus
Eb
1 -
Eb
2 -
Eb
3 in
δ configuration; tibial spurs present on legs III–IV; pedipalp chela finger median denticle rows non-imbricated; and posterior margins of tergite VII & metasoma I–III bearing fringes of microsetae. Within this group, it represents a sister genus of
Gint
KovařÍk et al., 2013
, where its
tYpe
species,
S. maidensis
comb
.
n
.
, was originallY placed despite several differences. Further DNA (
Just et al., 2022
) and phYlogenetic (KovařÍk & Lowe, 2022) analYses, however, clearlY demonstrated that
S. maidensis
comb
.
n
.
belongs to a separate, new genus. The two genera differ in the shape of the hemispermatophore, which has a large, tall, subtriangular hook-like basal lobe in
Sanaag
gen
.
n
.
and a small to medium sized, low, rounded, scoop-like basal lobe in
Gint
. Also, telson is rather bulbous in
Sanaag
gen
.
n
.
(telson L/D ratio
2.67–2.78 in
males) and rather elongated in
Gint
(telson L/D ratio
2.98–3.60 in
males).