The maculosus group of the genus Celaenorrhinus Hübner, 1819 from China, with discovery of female C. kuznetsovi Devyatkin, 2000 (Lepidoptera: Hesperiidae: Pyrginae)
Author
Huang, Zhen-Fu
Author
Chiba, Hideyuki
Author
Fei, Wen
Author
Fan, Xiao-Ling
text
Zootaxa
2014
3881
3
279
290
journal article
42273
10.11646/zootaxa.3881.3.6
b31f7509-f022-4a9e-9eda-0373fea229c6
1175-5326
230199
C1E7F95E-B3FC-4D14-A46E-A93B1556E664
Celaenorrhinus maculosus
(Felder & Felder, 1867)
Figs. 1–4
,
28–34
Pterygospidea maculosa
Felder & Felder, 1867: 528
, pl. 73, fig. 7 (
Type
locality: Shanghai).
Celaenorrhinus maculosa
: Watson, 1893: 49
; Leech, 1893: 569, pl. 39, fig. 2.
Celaenorrhinus maculosa maculosa
:
Evans, 1949
: 93
.
Celaenorrhinus maculosus maculosus
: Tong
(ed.), 1993: 66, pl. 58, figs. 635–636;
Chou, 1994
: 703
; Huang (ed.), 2001: 133.
Diagnosis.
Forewing length
22–25 mm
. Wings brown. Forewing upperside with inner margin of subapical spots in R3–R5 almost in line; spot in space M3 smaller than spots in M1, M2 and CuA1; spots in cell and space CuA1 separated apart; two spots in CuA2 separated, lower spot larger; forewing underside with basal yellow striping well developed. Hindwing upperside with the spot in space Rs large and distinct.
FIGURES 1–20
. Adult of the
masulosus
group species: 1–4.
C. maculosus
(Felder & Felder) (Guangdong)
, male (1–2), female (3–4); 5–6.
C. taiwanus
Matsumura
, male (Taiwan); 7–8.
C. osculus
Evens
, male (Guangdong); 9–12.
C. major
Hsu (Taiwan)
, male (9–10), female (11–12); 13–16.
C. inexpectus
Devyatkin (Guangdong)
, male (13–14), female (15–16); 17–24.
C. kuznetsovi
Devyatkin
, male (Guangxi) (17–18, 21–22), female (19–20 (Guangdong); 23–24 (Guangxi)); 25–27.
C. yaojiani
Huang & Wu
(Guangxi, holotype). Odd numbers, upperside; even numbers, underside; scale bar, 10 mm.
FIGURES 28–33
. Male genitalia of
C. maculosus
(Felder & Felder) (Guangdong)
. 28. ring, lateral view; 29. tegumen, ventral view; 30. right valva, inner view; 31–32. aedeagus; 33. juxta.
FIGURE 34
. Female genitalia of
C. maculosus
(Felder & Felder) (Guangdong)
.
Male genitalia (
Figs. 28–33
). Gnathos densely covered with spines and truncated end; valva nearly rectangular with dorso-distal part small and V-shaped; aedeagus robust, slightly longer than valva, with a curved cornuti, suprazonal sheath longer than subzonal sheath; juxta quadrifurcated with lateral branches shorter than half central branches, and with spherical mid-basal process.
Female genitalia (
Fig. 34
). Papillae anales triangular, covered with hairs; posterior apophyses slender, longer than papillae; posterior lamella broad, weakly sclerotized, with almost rounded or rectangular distal projection; ductus bursae screlotized with middle part narrow and distal part expanded; bursa copulatrix membranous, elongate.
Materials examined.
1♂
, Heishiding Nature Reserve, Guangdong,
14-V-1997
,
WANG
Min;
2♂
, Nanling National Nature Reserve, Guangdong,
22-VI-2003
,
20-V-1997
,
WANG
Min;
1♂
, Chebaling National Nature Reserve, Guangdong,
14-VIII-1989
,
WANG
Min; 1♀, Yaan, Sichuan,
FAN
Xiao-Ling,
7-VII-2003
; 1♀, Dashahe, Guizhou,
14-VIII-2011
, XU Hai-Ming. 1♀, Nanling National Nature Reserve, Guangdong,
29-VI-2012
,
DENG
Xiao-Hua, XU Hai-Ming;
1♂
Qingliangfeng National Nature Reserve, Zhejiang,
6-VIII-2012
, XU Hai-Ming.
Distribution
(
Fig. 68
).
China
(Sichuan, Henan, Shanghai, Jiangsu, Hubei, Zhejiang, Fujian, Guangdong),
Laos
,
Vietnam
.
Remarks.
The species, described by Felder & Felder (1867) from Shanghai, is widely distributed, and there are some geographic differences within
China
(
Devyatkin, 2000
). We compared the specimens from different localities. Wing markings and male genitalia are identical but the forewing length shows a slight difference. Specimens from South West
China
(female, n=
2, 22–23 mm
) and South
China
(male, n=
4, 22–24 mm
and female, n=
1, 22mm
) are smaller than that from East
China
(Zhejiang) near the
type
locality (male, n=
1, 25 mm
).