Short-legged daddy-long-leg spiders in North America: the genera Pholcophora and Tolteca (Araneae, Pholcidae) Author Huber, Bernhard A. 33607F65-19BF-4DC9-94FD-4BB88CED455F Zoological Research Museum Alexander Koenig, LIB, Bonn, Germany. Laboratory of Arachnology (LATLAX), Institute of Biology, Universidad Nacional Autónoma de México-Tlaxcala, San Miguel Contla, Santa Cruz Tlaxcala, Tlaxcala, México. Department of Genetics and Microbiology, Faculty of Science, Charles University, Prague, Czech Republic. Campus Amílcar Ferreira Sobral, Universidade Federal do Piauí, Floriano, Piauí, Brazil. b.huber@leibniz-lib.de Author Meng, Guanliang 7E8C41F8-77BB-468F-BE9A-D3F1DFCA1E4E Zoological Research Museum Alexander Koenig, LIB, Bonn, Germany. Laboratory of Arachnology (LATLAX), Institute of Biology, Universidad Nacional Autónoma de México-Tlaxcala, San Miguel Contla, Santa Cruz Tlaxcala, Tlaxcala, México. Department of Genetics and Microbiology, Faculty of Science, Charles University, Prague, Czech Republic. Campus Amílcar Ferreira Sobral, Universidade Federal do Piauí, Floriano, Piauí, Brazil. G.Meng@leibniz-lib.de Author Valdez-Mondragón, Alejandro F043A1C7-2B83-40C9-A74E-82C92F00725A lat_mactans@yahoo.com.mx Author Král, Jiří E836F3B5-D704-4EEC-966A-0C4F1FAD324B spider@natur.cuni.cz Author Ávila Herrera, Ivalú M. E3687584-7F64-450D-9492-BE0DD4864AD6 ivalu.a@gmail.com Author Carvalho, Leonardo S. 28AA7D67-3C9D-495E-8C17-33D35F1A0FAC carvalho@ufpi.edu.br text European Journal of Taxonomy 2023 2023-07-17 880 1 1 89 http://zoobank.org/3f806fd6-2eb3-456a-afd7-780a0fbeb2da journal article 55344 10.5852/ejt.2023.880.2173 339bce42-a7e7-418b-b9ac-4588e4219b15 2118-9773 8155472 3F806FD6-2EB3-456A-AFD7-780A0FBEB2DA Pholcophora texana Gertsch, 1935 Figs 3H–I , 19–24 , 33I–J Remark For synonymy, type material, and redescription, see Huber (2000) . Diagnosis Easily distinguished from known congeners by unique brush of modified hairs on male palpal femur ( Fig. 19A, C ) and by pair of round sacs in female internal genitalia ( Fig. 22B–D ); also by shape of procursus ( Fig. 20C–E ; tip in lateral view with wide transparent flap), by shape of distal bulbal process ( Fig. 20F–H ), and by indistinct plate in front of main epigynal plate ( Fig. 21A, C ). Material examined (new record) MEXICO – Hidalgo • 1 ♂ , 1 ♀ , 3 female abdomens; ~ 2.5 km SW of Jacala ; 20.9948° N , 99.2138° W ; 1430 m a.s.l. ; 21 Oct. 2019 ; B.A. Huber and A. Valdez-Mondragón leg.; ZFMK Ar 23952 • 7 ♀♀ , 2 juvs, in pure ethanol ( three female prosomata used for molecular work, abdomens transferred to ZFMK Ar 23952); same collection data as for preceding; ZFMK Mex341 • 6 ♀♀ , 2 juvs (subadult ♂♂ ); same collection data as for preceding; LATLAX . Description (amendments; see Huber 2000 ) Male Measurements of male from SW of Jacala, Hidalgo (ZFMK Ar 23952): body length 1.75, carapace width 0.70; tibia 1 length 1.25, metatarsus 1 length 1.30, tibia 2 length 0.95, tibia 4 length 1.25. Distance PME-PME 70 µm; diameter PME 50 µm; distance PME-ALE 25 µm; distance AME-AME 15 µm; diameter AME 30 µm. Diameters of leg femora 0.14–0.16, of leg tibiae 0.08–0.09. Clypeus unmodified, but slightly more protruding than in female; clypeus rim to ALE 0.21. Distances between cheliceral stridulatory ridges proximally 2.0 µm, distally 3.2 µm. Sternum humps very distinct, ~0.08 long. Procursus as in Fig. 20C–E , with distinctive transparent element distally (barely visible in dissecting microscope); genital bulb as in Fig. 20F–H . Leg tibiae with slightly higher number of vertical hairs than in female (proximally on tibiae 1 and 2); prolateral trichobothrium absent on tibia 1. Tibia 1 in previously measured specimens: 0.72 ( Gertsch 1982 ); 0.96, 0.97, 1.39 ( Huber 2000 ). Female In general, similar to male but sternum without pair of anterior humps, and chelicerae without stridulatory files. Tibia 1 in 11 females from SW of Jacala: 0.90–1.18 (mean 1.03). Epigynum and internal female genitalia as in Figs 21 , 22 , 23C , with pair of receptacles (shape varies slightly with angle of view), without (or with very small?) pore plates. Fig. 19. Pholcophora texana Gertsch, 1935 , male from Mexico, Hidalgo, SW of Jacala (ZFMK Ar 23952). Left palp, prolateral, dorsal, and retrolateral views; arrows point at unique brush of modified hairs. Scale bar = 0.3 mm. Distribution Apparently widely distributed in north-eastern Mexico , ranging into southern Texas ( Fig. 2 ). However, the map in Fig. 2 includes four records that are based exclusively on females, i.e., the identifications should be verified (e.g., by collecting males or by sequencing specimens from these localities). Natural history The newly collected specimens were found under rocks on an arid hillside set with low thorn scrub ( Fig. 34B ).