Revision of the genera Heteroptilon, Nugonioneura and Opistocladus from the Lower Permian of USA (Insecta: Cnemidolestida: Tillyardembiidae, Nugonioneuridae and Parmapteridae) Author Aristov, D. S. text Far Eastern Entomologist 2020 2020-02-04 401 1 9 http://dx.doi.org/10.25221/fee.401.1 journal article 10.25221/fee.401.1 2713-2196 7165429 B48F9993-2A8E-4749-BCFD-C1211D0F4B47 Heteroptilon costale Carpenter, 1976 Figs 1, 2 Heteroptilon costale Carpenter, 1976: 347, fig. 8; 1992: 126, fig. 73, 7; Béthoux et al ., 2003: 278. MATERIAL. Holotype MCZ , № 5878, positive imprint of deformed forewing; United States , Kansas , Dickinson County, Banner Township, 5 km southeast of Elmo town, Elmo locality; Lower Permian, Lower Leonardian, Sumner Group, Wellington Formation, Carlton Member; in MCZ . REDESCRIPTION. Anterior margin of forewing is convex. Anterior branches of SC near wings mid length are numerous and long, strongly inclined anteriorly. RS branching early and abundantly, posteriorly subpectinate, with nine branches. Anterior branch of M simple, posterior with two apices. CuA with three branches. А 1 simple, А 2 bifurcated. Transverse veins simple, numerous in subcostal field. MEASUREMENTS. Forewing length 15 mm .