Taxonomy and evolution of asymmetric male genitalia in the subgenus Ashima Chen (Diptera: Drosophilidae: Phortica Schiner), with descriptions of seven new species Author Toda, Masanori J. Hokkaido University Museum, Hokkaido University, Sapporo 060 - 0810, Japan Author Bänziger, Hans Department of Entomology and Plant Pathology, Faculty of Agriculture, Chiang Mai University, Chiang Mai 50200, Thailand hans. banziger @ cmu. ac. th; https: // orcid. org / 0000 - 0001 - 8913 - 3034 Author Sati, Pradeep C. Systematic & Cytogenetic Laboratory, Department of Zoology and Biotechnology, HNB Garhwal University, Srinagar-Garhwal, 246174, Uttarakhand, India. & satipradeep 28 @ gmail. com; https: // orcid. org / 0000 - 0001 - 6809 - 8575 Author Fartyal, Rajendra S. Systematic & Cytogenetic Laboratory, Department of Zoology and Biotechnology, HNB Garhwal University, Srinagar-Garhwal, 246174, Uttarakhand, India. & rs. fartyal @ hnbgu. ac. in; https: // orcid. org / 0000 - 0002 - 8844 - 6263 Author Suwito, Awit Zoology Division (Museum Zoologicum Bogoriense), Research Center for Biology-LIPI, Jl. Raya Jakarta-Bogor Km. 46, Cibinong 16911, West Java, Indonesia. awitswt @ yahoo. com Author Katoh, Toru Department of Biological Sciences, Faculty of Science, Hokkaido University, Sapporo 060 - 0810, Japan t _ katoh @ sci. hokudai. ac. jp; https: // orcid. org / 0000 - 0003 - 3376 - 8545 text Zootaxa 2020 2020-06-08 4789 1 1 54 journal article 21673 10.11646/zootaxa.4789.1.1 c12aaed8-264c-459c-b1aa-09eaeb686670 1175-5326 3884610 311A27A2-A053-4D20-A43E-EF57DBE55166 30) Phortica ( Ashima ) idiasta Gong, Zhu & Chen, 2018 Phortica ( Ashima ) idiasta Gong, Zhu & Chen, 2018: 440 . Diagnosis. Pregonite partly less sclerotized (ch.43-0), not serrated on ventrodistal portion (ch.50-0); ventrosubapical portion of apical knob more or less protruded, forming barbule (ch.46-0); posterior postgonite dorsad curved, claw-like process, and anterior one basally divided into 3 long, outward oriented processes (“Figs 24, 25” in Gong et al . 2018 ). Supplementary description. Supracervical setae approximately 14. Dorsomedial, tentorial apodeme 1/2 as long as basal, parallel portion of dorsolateral, tentorial apodeme. Arista with arb = 2/0; longest, dorsal branch as long as longest seta on pedicel; apical, leaf-like expansion narrower than long, apically roundish and with minute projection. Cibarial, medial sensilla 6–7 per side; posterior sensilla 3–4 per side. The antisymmetry is observed in the phallic organs: in A-type, the posterior postgonite is present on the right side, the anterior one on the left side, and left gonocoxite of hypandrium more stretched out (“Fig. 25” in Gong et al . 2018 ); but in B-type, vice versa . Specimen examined. Thailand : 1♂ (B-type), Umphang , Tak , 30.v.2008 , H. Bänziger leg. ( SEHU ) . Distribution. China ( Yunnan ), Thailand *. Remarks. This species first branched off in the glabra subgroup ( Fig. 1 ). The remaining three species form a cluster (BP = 96%) with three synapomorphies ( Fig. 1 ): pregonite ( Fig. 9G ) nearly entirely sclerotized (ch.43-1), only angulated in ventrosubapical portion of apical knob (ch.46-1), and coarsely serrated on ventrodistal portion (ch.50-1).