Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae) Author Ballantyne, Lesley A. Author Lambkin, Christine text Zootaxa 2009 2009-02-04 1997 1 188 journal article 1175­5334 Atyphella atra Lea, 1921 ( Figs 9, 10 , 83–85 ) Atyphella atra Lea, 1921a:198 . Ballantyne in Calder 1998: 177 (comb. rev.) Ballantyne & Lambkin, 2000: 25 ; 2001: 374, 2006: 30. Luciola (Luciola) atra (Lea) . McDermott, 1966: 99 . Luciola (Atyphella) atra (Lea) . Ballantyne & McLean, 1970: 281 . Moore, 1989: 136 . Holotype . Male. AUSTRALIA : 28.19S , 153.05E , Lamington National Park (C2290 QM ). Diagnosis. Males distinguished from most other Atyphella by the acute frons-vertex junction, the small (probably non–functional) mouthparts, the contiguous antennal sockets, the brown – dark brown parallelsided elytra; females pale coloured with fully developed elytra and vestigial hind wings; paler male specimens very like A. similis , which has angulate posterolateral pronotal corners, separated antennal sockets, well developed mouthparts and flightless brachelytral females. Ballantyne and Lambkin (2000) distinguished atra and similis most obviously by their distinctive females.