Fregetta lineata (Peale, 1848) is a valid extant species endemic to New Caledonia
Author
Bretagnolle, Vincent
Author
Flood, Robert L.
Author
Gaba, Sabrina
Author
Shirihai, Hadoram
text
Bulletin of the British Ornithologists’ Club
2022
2022-03-11
142
1
111
130
https://bioone.org/journals/bulletin-of-the-british-ornithologists-club/volume-142/issue-1/bboc.v142i1.2022.a6/Fregetta-lineata-Peale-1848-is-a-valid-extant-species-endemic/10.25226/bboc.v142i1.2022.a6.full
journal article
10.25226/bboc.v142i1.2022.a6
2513-9894
12535700
B48F3D74-2BFE-49A1-A560-C5E59CF8FD18
Phylogenetic relationships between
Fregetta
storm petrels
Recent genetic work shed light on the
Fregetta
complex, but also added uncertainties and confusion. These have confirmed that live
F. maoriana
and the
three
F. maoriana
specimens are the same taxon (
Robertson
et al
. 2011
). They are distinct from and not close to
Oceanites
and lie within
Fregetta
(
Robertson
et al
. 2011
,
2016
). This is supported by evidence from Mallophaga:
Philoceanus fasciatus
, a species found on both
F. tropica
and
F. grallaria
, has been collected on
F. maoriana
, and is morphologically very different from both
Philoceanus robertsi
, parasitic on
Oceanites oceanicus
, and
Philoceanus garrodiae
, parasitic on Grey-backed Storm Petrel
Garrodia nereis
(
Stephenson
et al
. 2008b
)
.
Figure 6. Comparative measurements of New Caledonian Storm Petrel
Fregetta lineata
(black), White-bellied Storm Petrel
F. grallaria
(red) and Black-bellied Storm Petrel
F. tropica
(green). Data are presented as a correlation matrix for each pair of morphometric variables, with the diagonal in the matrix showing the distribution of each variable per taxon. Colour chart as in Fig. 4F.
Three studies provide
Fregetta
phylogenetic trees; based on either short (
c
.500 bp;
Cibois
et al
. 2015
), long (960 bp;
Robertson
et al
. 2016
), or complete cyt-
b
sequences (
Robertson
et al
. 2011
); as well as seventh intron of beta Fib (
Robertson
et al
. 2011
,
2016
). This diversity of genetic material led to contrasting conclusions.
F. maoriana
was closer to
F. tropica
than
F. grallaria
in both complete cyt-
b
and nuclear intron (
Robertson
et al
. 2011
), closer to
F. tropica
than
F. grallaria
in short cyt-
b
(
Cibois
et al
. 2015
), but closer to
F. grallaria
in long cyt-
b
and closer to
F. tropica
in nuclear intron (
Robertson
et al
. 2016
). In addition,
F. lineata
AMNH
194110 was sequenced and results indicate that it diverges from
F. maoriana
and, based on partial cyt-
b
sequence (557 bp), is more closely related to
F. grallaria
, especially
F.
[
g.
]
titan
, than to
F. tropica
(
Cibois
et al
. 2015
,
Robertson
et al
. 2016
). A matter of note,
Robertson
et al
. (2016)
commented that
Cibois
et al
. (2015)
were wrong to include
NHMUK
1953.55.101 (collected on Gough Island, South Atlantic, held at the Natural History Museum, Tring) as
F. grallaria
. However, the sequence was taken from
Robertson
et al
. (2011)
, as recorded in GenBank, so the original error is owned by the latter.
Our findings show that
USNM
15713 and
AMNH
194110 are the same taxon,
F. lineata
, based on morphometrics, although no genetic data are available for the
USNM
specimen. Interestingly, the Brisbane specimen
QM
14391 was sequenced and clumped with
F.
[
g.
]
titan
(
Robertson
et al
. 2011
), as did the Ua Pou specimen
AMNH
194110 (
Cibois
et al
. 2015
), supporting the case for aggregating them. Incidentally, whilst
QM
14391 is labelled
F. tropica
,
Robertson
et al
. (2016)
treated it as
F. grallaria
, presumably because sequences clade with
F. grallaria
rather than
F. tropica
. However, systematics of the group are complicated by the opaque taxonomy of the genus
Fregetta
itself (
Howell 2010
,
2012
), with more than 20 different names proposed for these birds, and the complex situation in
Tristan da Cunha
, South Atlantic (
Brooke 2004
,
Howell 2012
,
Flood & Fisher 2013
). In particular, taxonomy of the
F. grallaria
and
F. tropica
complexes are not resolved (
Crochet 2008
,
Howell 2010
,
2012
,
Robertson
et al
. 2016
). That said, there is consensus that the four
Fregetta
taxa (
F. grallaria
,
F. tropica
,
F. lineata
and
F. maoriana
) form a monophyletic clade, based on mtDNA and nuclear DNA, albeit a single gene in both cases (
Robertson
et al
. 2011
,
Cibois
et al
. 2015
,
Robertson
et al
. 2016
). This clade is distinct from
Oceanites
and supports the generic denomination
Fregetta
.