The beetle family Carabidae of Costa Rica: The genera of the Cryptobatida group of subtribe Agrina, tribe Lebiini, with new species and notes on their way of life (Insecta: Coleoptera) Author Erwin, Terry L. text Zootaxa 2004 662 1 54 journal article 10.5281/zenodo.158702 64a4a578-1107-4369-b2c4-67921dd0664c 1175­5326 158702 E1878D62-01DE-4BB2-8960-23085056D27F Otoglossa Chaudoir 1872 Otoglossa Chaudoir 1872 :158 . Type species: Otoglossa tuberculosa Chaudoir 1872 :158 (designated by Reichardt 1964 :49 ). Type locality: BRAZIL , Minas Gerias. Heraldinium Liebke, 1927 :101 . Type species: Heraldinium nevermanni Liebke, 1927 :102 , (original designation). Synonymized by Reichardt 1964 :49 . Diagnosis. (cf. Fig. 23) Form unusual for lebiine carabids: head and pronotum markedly narrow, elytra broad and box­like with multiple tubercles or calluses on intervals 1–7. Prothorax nearly cylindrical, with very narrow and slightly dentate lateral pronotal bead. Head with large semi­hemispheric eyes and gradually, yet markedly, constricted neck. Male and female with small v­shaped indentation medial on posterior margin of sternum 6. Male endophallus with flagellum, flagellum apically barbed. Geographic distribution. The known range of this widespread Neotropical genus extends from Costa Rica south into southeastern Perú , east to French Guiana and south to southeastern Brasil (Rio de Janeiro). Notes. This genus is now known to contain three species. Amongst the Cryptobatia Group, only male adults of Otoglossa and Pseudotoglossa have a barbed apex on the flagellum of the endophallus of the male genitalia. Whilst the adults have a very different aspect, it must be noted here that a rigorous phylogenetic analysis may indicate that the two genera should be classified in two subgenera within the genus Otoglossa .