<document id="7B31329D3D34E29E6699CC2F98EC56E4" ID-CLB-Dataset="29485" ID-DOI="10.11646/zootaxa.4471.2.1" ID-GBIF-Dataset="bb5c58a5-3a4a-4c8b-8025-7e71e77c2782" ID-ISSN="1175-5326" ID-Zenodo-Dep="1439596" ID-ZooBank="C2615AC4-83EF-46F0-8943-85B42A03745C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538363978247" checkinUser="plazi" docAuthor="Smetana, Aleš" docDate="2018" docId="0A2E0E6AF906FFC1FF46FD4CFC21FA55" docLanguage="en" docName="zootaxa.4471.2.1.pdf" docOrigin="Zootaxa 4471 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Agelosus sikkimensis Bernhauer 1920" docType="treatment" docVersion="8" lastPageNumber="224" masterDocId="F6177612F911FFD6FFD1FFC6FFC4FF80" masterDocTitle="Review of the genera Agelosus Sharp, 1889, Apostenolinus Bernhauer, 1934 and Apecholinus Bernhauer, 1933 (Coleoptera: Staphylinidae: Staphylinini: Staphylinina)" masterLastPageNumber="244" masterPageNumber="201" pageNumber="224" updateTime="1698558130547" updateUser="ExternalLinkService">
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<mods:title id="7C182ACF8D1EFF790320A7F1814454C1">Review of the genera Agelosus Sharp, 1889, Apostenolinus Bernhauer, 1934 and Apecholinus Bernhauer, 1933 (Coleoptera: Staphylinidae: Staphylinini: Staphylinina)</mods:title>
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<taxonomicName id="4587C4FFF906FFC1FF46FD4CFDA9FD24" ID-CoL="5TPY8" authority="Bernhauer, 1920" authorityName="Bernhauer" authorityYear="1920" box="[151,621,650,676]" class="Insecta" family="Staphylinidae" genus="Agelosus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="sikkimensis">
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<emphasis id="B0F3636EF906FFC1FF46FD4CFE53FD24" bold="true" box="[151,407,650,676]" italics="true" pageId="23" pageNumber="224">Agelosus sikkimensis</emphasis>
<bibRefCitation id="E616C28DF906FFC1FE4FFD4CFDA9FD24" box="[414,621,650,676]" pageId="23" pageNumber="224" refId="ref31990" refString="Bernhauer, M. (1920) Neue Staphyliniden des indo-malaischen Gebietes. Archiv fur Naturgeschichte (A), 84, 177 - 188." type="journal article">Bernhauer, 1920</bibRefCitation>
</emphasis>
</taxonomicName>
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(
<figureCitation id="1ABCA3F9F906FFC1FF4EFD6BFF37FD46" box="[159,243,685,710]" captionStart="FIGURES 41–48. 41–46" captionStartId="20.[151,264,1890,1912]" captionTargetBox="[296,1314,317,1718]" captionTargetId="figure@20.[150,1437,195,1868]" captionTargetPageId="20" captionText="FIGURES 41–48. 41–46. Agelosus quadrimaculatus: 41,43, tergite 10 of male genital segment; 42, sternite 9 of male genital segment; 44, apical portion of sternite 9 of male genital segment; 45,46, apical portions of tergite 10 of female genital segment. 47–48. Agelosus sikkimensis: 47, aedoeagus, ventral view; 48, apical portion of underside of paramere with sensory peg setae." httpUri="https://zenodo.org/record/1439602/files/figure.png" pageId="23" pageNumber="224">Figs 47</figureCitation>
–50,83–87)
</paragraph>
</subSubSection>
<subSubSection id="CA9DECF7F906FFC1FF46FD36FA58FCA4" pageId="23" pageNumber="224" type="reference_group">
<paragraph id="8238BF7CF906FFC1FF46FD36FA5FFC86" blockId="23.[151,1437,752,1142]" box="[151,1435,752,774]" pageId="23" pageNumber="224">
<treatmentCitationGroup id="A2979852F906FFC1FF46FD36FA5FFC86" box="[151,1435,752,774]" pageId="23" pageNumber="224">
<taxonomicName id="4587C4FFF906FFC1FF46FD36FDC8FC86" ID-CoL="5TPY8" authority="Bernhauer, 1920: 185" authorityName="Bernhauer" authorityPageNumber="185" authorityYear="1920" box="[151,524,752,774]" class="Insecta" family="Staphylinidae" genus="Agelosus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="sikkimensis">
<emphasis id="B0F3636EF906FFC1FF46FD36FECEFC86" box="[151,266,752,774]" italics="true" pageId="23" pageNumber="224">sikkimensis</emphasis>
<treatmentCitation id="0326996DF906FFC1FECAFD36FDC8FC86" author="" box="[283,524,752,774]" page="185" pageId="23" pageNumber="224">
<bibRefCitation id="E616C28DF906FFC1FECAFD36FE15FC86" box="[283,465,752,774]" pageId="23" pageNumber="224" refId="ref31990" refString="Bernhauer, M. (1920) Neue Staphyliniden des indo-malaischen Gebietes. Archiv fur Naturgeschichte (A), 84, 177 - 188." type="journal article">Bernhauer, 1920</bibRefCitation>
: 185
</treatmentCitation>
</taxonomicName>
(
<taxonomicName id="4587C4FFF906FFC1FDF4FD36FCCFFC86" ID-CoL="8VS43" authority="(Tasgius)" authorityName="Tasgius" box="[549,779,752,774]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="B0F3636EF906FFC1FDF4FD36FD5EFC86" box="[549,666,752,774]" italics="true" pageId="23" pageNumber="224">Staphylinus</emphasis>
(
<emphasis id="B0F3636EF906FFC1FD65FD37FCC4FC86" box="[692,768,753,774]" italics="true" pageId="23" pageNumber="224">Tasgius</emphasis>
)
</taxonomicName>
; description); Cameron, 1932: 206 (
<taxonomicName id="4587C4FFF906FFC1FB60FD36FA52FC86" ID-CoL="8VS43" authority="(Tasgius)" authorityName="Tasgius" box="[1201,1430,752,774]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="B0F3636EF906FFC1FB60FD36FAE2FC86" box="[1201,1318,752,774]" italics="true" pageId="23" pageNumber="224">Staphylinus</emphasis>
(
<emphasis id="B0F3636EF906FFC1FA91FD37FA48FC86" box="[1344,1420,753,774]" italics="true" pageId="23" pageNumber="224">Tasgius</emphasis>
)
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;
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redescription);) 
<bibRefCitation id="E616C28DF906FFC1FEACFCC8FDF9FCA4" box="[381,573,782,804]" pageId="23" pageNumber="224" refId="ref33499" refString="Scheerpeltz, O. (1933) Staphylinidae VII (Pars 129). Supplementum I. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum Catalogus. Vol. VI. Staphylinidae. Junk, Berlin, pp. 989 - 1500." type="book chapter">Scheerpeltz, 1933</bibRefCitation>
: 1402 (
<taxonomicName id="4587C4FFF906FFC1FD47FCC8FCCFFCA4" ID-CoL="8VS43" box="[662,779,782,804]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="B0F3636EF906FFC1FD47FCC8FCCFFCA4" box="[662,779,782,804]" italics="true" pageId="23" pageNumber="224">Staphylinus</emphasis>
</taxonomicName>
; catalogue); 
<bibRefCitation id="E616C28DF906FFC1FC4CFCC8FBF1FCA4" box="[925,1077,782,804]" pageId="23" pageNumber="224" refId="ref32242" refString="Coiffait, H. (1977) Ergebnisse der Bhutan- Expedition 1972 des Naturhistorischen Museums in Basel. Coleoptera: Fam. Staphylinidae Subfam. Xantholininae et Staphylininae. Entomologica Basiliensia, 2, 205 - 242." type="journal article">Coiffait, 1977</bibRefCitation>
: 212, 
<figureCitation id="1ABCA3F9F906FFC1FBAEFCC8FB39FCA4" box="[1151,1277,782,804]" captionStart="FIGURES 1–13. 1–13" captionStartId="6.[151,264,1877,1899]" captionTargetBox="[240,1326,391,1749]" captionTargetId="figure@6.[150,1437,229,1902]" captionTargetPageId="6" captionText="FIGURES 1–13. 1–13. Agelosus carinatus: 1,3,4, aedoeagus, ventral view; 2,6,8,10–13, apical portions of underside of paramere with sensory peg setae; 5,7,9,11, apical portions of paramere, ventral view." httpUri="https://zenodo.org/record/1439598/files/figure.png" pageId="23" pageNumber="224">Figs 5 G,H</figureCitation>
; 211 (
<taxonomicName id="4587C4FFF906FFC1FA99FCD6FA50FCA5" ID-CoL="8VS53" box="[1352,1428,784,805]" class="Insecta" family="Staphylinidae" genus="Tasgius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="B0F3636EF906FFC1FA99FCD6FA50FCA5" box="[1352,1428,784,805]" italics="true" pageId="23" pageNumber="224">Tasgius</emphasis>
</taxonomicName>
;
</paragraph>
</subSubSection>
<subSubSection id="CA9DECF7F906FFC1FF16FCEBFD6AFBF6" pageId="23" pageNumber="224" type="description">
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aedoeagus illustrated; 
<collectingCountry id="FA90FFECF906FFC1FE78FCEBFE36FCC3" box="[425,498,813,835]" name="Bhutan" pageId="23" pageNumber="224">Bhutan</collectingCountry>
record); 
<bibRefCitation id="E616C28DF906FFC1FD9EFCE8FD26FCC3" box="[591,738,813,835]" pageId="23" pageNumber="224" refId="ref32612" refString="Herman, L. H. (2001) Catalogue of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. VI. Staphylinine group. Part 3. Staphylininae: Staphylinini (Quediina, Staphylinina, Tanygnathinina, Xanthopygina). Xantholinini. Staphylinidae incertae sedis, fossils. Protactinae. Bulletin of the American Museum of Natural History, 265, 3021 - 3840." type="journal article">Herman, 2001</bibRefCitation>
:3550 (
<taxonomicName id="4587C4FFF906FFC1FCF8FCE8FCB1FCC3" authorityName="Smetana" authorityYear="2018" box="[809,885,814,835]" class="Insecta" family="Staphylinidae" genus="Ocypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="B0F3636EF906FFC1FCF8FCE8FCB1FCC3" box="[809,885,814,835]" italics="true" pageId="23" pageNumber="224">Ocypus</emphasis>
</taxonomicName>
; catalogue); 
<bibRefCitation id="E616C28DF906FFC1FC28FCEBFB50FCC3" box="[1017,1172,813,835]" pageId="23" pageNumber="224" refId="ref34017" refString="Smetana, A, (2004) Staphylininae. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Apollo Books, Stenstrup, pp. 624 - 698." type="book chapter">Smetana, 2004</bibRefCitation>
: 676 (
<taxonomicName id="4587C4FFF906FFC1FB03FCE8FADAFCC3" authorityName="Smetana" authorityYear="2018" box="[1234,1310,814,835]" class="Insecta" family="Staphylinidae" genus="Ocypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="B0F3636EF906FFC1FB03FCE8FADAFCC3" box="[1234,1310,814,835]" italics="true" pageId="23" pageNumber="224">Ocypus</emphasis>
</taxonomicName>
; catalogue);
</paragraph>
<paragraph id="8238BF7CF906FFC1FF16FC8AFA58FC00" blockId="23.[151,1437,752,1142]" pageId="23" pageNumber="224">
<bibRefCitation id="E616C28DF906FFC1FF16FC8AFE16FCE2" box="[199,466,844,866]" pageId="23" pageNumber="224" refId="ref33592" refString="Schulke, M. &amp; Smetana, A. (2015) Staphylinina. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Revised and updated Edition. Brill, Leiden, pp. 1081 - 1107." type="book chapter">Schülke &amp; Smetana, 2015</bibRefCitation>
: 1090 (
<taxonomicName id="4587C4FFF906FFC1FDCFFC8AFDAEFCE1" authorityName="Smetana" authorityYear="2018" box="[542,618,844,865]" class="Insecta" family="Staphylinidae" genus="Ocypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="B0F3636EF906FFC1FDCFFC8AFDAEFCE1" box="[542,618,844,865]" italics="true" pageId="23" pageNumber="224">Ocypus</emphasis>
</taxonomicName>
, catalogue) 
<taxonomicName id="4587C4FFF906FFC1FF46FCADFDC5FC00" authority="Cameron, 1932: 206" authorityName="Cameron" authorityPageNumber="206" authorityYear="1932" box="[151,513,874,896]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="auroguttatus">
<emphasis id="B0F3636EF906FFC1FF46FCADFEDCFC00" box="[151,280,875,896]" italics="true" pageId="23" pageNumber="224">auroguttatus</emphasis>
Cameron, 1932: 206
</taxonomicName>
(
<taxonomicName id="4587C4FFF906FFC1FDC7FCACFD33FC00" authority="(Tasgius)" authorityName="Tasgius" box="[534,759,874,896]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="B0F3636EF906FFC1FDC7FCACFD4FFC00" box="[534,651,874,896]" italics="true" pageId="23" pageNumber="224">Staphylinus</emphasis>
(
<emphasis id="B0F3636EF906FFC1FD70FCADFD29FC00" box="[673,749,875,896]" italics="true" pageId="23" pageNumber="224">Tasgius</emphasis>
)
</taxonomicName>
; description); 
<bibRefCitation id="E616C28DF906FFC1FC46FCACFB91FC00" box="[919,1109,874,896]" pageId="23" pageNumber="224" refId="ref33499" refString="Scheerpeltz, O. (1933) Staphylinidae VII (Pars 129). Supplementum I. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum Catalogus. Vol. VI. Staphylinidae. Junk, Berlin, pp. 989 - 1500." type="book chapter">Scheerpeltz, 1933</bibRefCitation>
:1391 (
<taxonomicName id="4587C4FFF906FFC1FB73FCACFAD3FC00" box="[1186,1303,874,896]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="B0F3636EF906FFC1FB73FCACFAD3FC00" box="[1186,1303,874,896]" italics="true" pageId="23" pageNumber="224">Staphylinus</emphasis>
</taxonomicName>
; catalogue);
</paragraph>
<paragraph id="8238BF7CF906FFC1FF16FC4FFA58FC1F" blockId="23.[151,1437,752,1142]" box="[199,1436,905,927]" pageId="23" pageNumber="224">
<bibRefCitation id="E616C28DF906FFC1FF16FC4FFEA6FC1F" box="[199,354,905,927]" pageId="23" pageNumber="224" refId="ref32280" refString="Coiffait, H. (1982 a) Staphylinides (Col.) de la region himalayenne et de l'Inde (I. Xantholininae, Staphylininae et Paederinae). Entomologica Basiliensia, 7, 231 - 302." type="journal article">Coiffait, 1982a</bibRefCitation>
:233 (
<taxonomicName id="4587C4FFF906FFC1FE4DFC4FFDD7FC1F" authorityName="Coiffait" authorityYear="1982" box="[412,531,905,927]" class="Insecta" family="Staphylinidae" genus="Nudabemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="B0F3636EF906FFC1FE4DFC4FFDD7FC1F" box="[412,531,905,927]" italics="true" pageId="23" pageNumber="224">Nudabemus</emphasis>
</taxonomicName>
; 
<collectingCountry id="FA90FFECF906FFC1FDF0FC4FFD99FC1F" box="[545,605,905,927]" name="Nepal" pageId="23" pageNumber="224">Nepal</collectingCountry>
and Darjeeling records); 
<bibRefCitation id="E616C28DF906FFC1FCB0FC4FFBA7FC1F" box="[865,1123,905,927]" pageId="23" pageNumber="224" refId="ref34111" refString="Smetana, A. &amp; Davies A. (2000) Reclassification of the north temperate taxa associated with Staphylinus sensu lato, including comments on relevant subtribes of Staphylinini (Coleoptera: Staphylinidae). American Museum Novitates, 3287, 1 - 88. https: // doi. org / 10.1206 / 0003 - 0082 (2000) 287 % 3 C 0001: ROTNTT % 3 E 2.0. CO; 2" type="journal article">Smetana &amp; Davies, 2000</bibRefCitation>
: 42 (
<taxonomicName id="4587C4FFF906FFC1FB44FC4CFB25FC1F" authorityName="Smetana" authorityYear="2018" box="[1173,1249,906,927]" class="Insecta" family="Staphylinidae" genus="Ocypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="B0F3636EF906FFC1FB44FC4CFB25FC1F" box="[1173,1249,906,927]" italics="true" pageId="23" pageNumber="224">Ocypus</emphasis>
</taxonomicName>
; listing); 
<bibRefCitation id="E616C28DF906FFC1FA95FC4CFA58FC1F" box="[1348,1436,906,927]" pageId="23" pageNumber="224" refId="ref32612" refString="Herman, L. H. (2001) Catalogue of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. VI. Staphylinine group. Part 3. Staphylininae: Staphylinini (Quediina, Staphylinina, Tanygnathinina, Xanthopygina). Xantholinini. Staphylinidae incertae sedis, fossils. Protactinae. Bulletin of the American Museum of Natural History, 265, 3021 - 3840." type="journal article">Herman,</bibRefCitation>
</paragraph>
<paragraph id="8238BF7CF906FFC1FF16FC6EFA5FFC3E" blockId="23.[151,1437,752,1142]" box="[199,1435,936,958]" pageId="23" pageNumber="224">
2001:3365 (
<taxonomicName id="4587C4FFF906FFC1FE97FC6FFE56FC3E" authorityName="Smetana" authorityYear="2018" box="[326,402,937,958]" class="Insecta" family="Staphylinidae" genus="Ocypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="B0F3636EF906FFC1FE97FC6FFE56FC3E" box="[326,402,937,958]" italics="true" pageId="23" pageNumber="224">Ocypus</emphasis>
</taxonomicName>
; catalogue); 
<bibRefCitation id="E616C28DF906FFC1FDCEFC6EFD7AFC3E" box="[543,702,936,958]" pageId="23" pageNumber="224" refId="ref34017" refString="Smetana, A, (2004) Staphylininae. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Apollo Books, Stenstrup, pp. 624 - 698." type="book chapter">Smetana, 2004</bibRefCitation>
: 676 (
<taxonomicName id="4587C4FFF906FFC1FCD5FC6FFC94FC3E" authorityName="Smetana" authorityYear="2018" box="[772,848,937,958]" class="Insecta" family="Staphylinidae" genus="Ocypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="B0F3636EF906FFC1FCD5FC6FFC94FC3E" box="[772,848,937,958]" italics="true" pageId="23" pageNumber="224">Ocypus</emphasis>
</taxonomicName>
; catalogue); 
<bibRefCitation id="E616C28DF906FFC1FC0CFC6EFB31FC3E" box="[989,1269,936,958]" pageId="23" pageNumber="224" refId="ref33592" refString="Schulke, M. &amp; Smetana, A. (2015) Staphylinina. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Revised and updated Edition. Brill, Leiden, pp. 1081 - 1107." type="book chapter">Schülke &amp; Smetana, 2015</bibRefCitation>
: 1089 (
<taxonomicName id="4587C4FFF906FFC1FA9BFC6FFA52FC3E" authorityName="Smetana" authorityYear="2018" box="[1354,1430,937,958]" class="Insecta" family="Staphylinidae" genus="Ocypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="B0F3636EF906FFC1FA9BFC6FFA52FC3E" box="[1354,1430,937,958]" italics="true" pageId="23" pageNumber="224">Ocypus</emphasis>
</taxonomicName>
,
</paragraph>
<paragraph id="8238BF7CF906FFC1FF16FC00FA58FC7B" blockId="23.[151,1437,752,1142]" pageId="23" pageNumber="224">
catalogue) 
<emphasis id="B0F3636EF906FFC1FEE7FC00FE6EFC5C" bold="true" box="[310,426,966,988]" pageId="23" pageNumber="224">comb. nov.</emphasis>
, 
<emphasis id="B0F3636EF906FFC1FE66FC00FDD0FC5C" bold="true" box="[439,532,966,988]" pageId="23" pageNumber="224">syn. nov.</emphasis>
<taxonomicName id="4587C4FFF906FFC1FF46FC23FE3AFC7B" authority="Cameron, 1932: 207" authorityName="Cameron" authorityPageNumber="207" authorityYear="1932" box="[151,510,997,1019]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="bimaculatus">
<emphasis id="B0F3636EF906FFC1FF46FC23FED7FC7B" box="[151,275,997,1019]" italics="true" pageId="23" pageNumber="224">bimaculatus</emphasis>
Cameron, 1932: 207
</taxonomicName>
(
<taxonomicName id="4587C4FFF906FFC1FDC5FC23FD31FC7B" authority="(Tasgius)" authorityName="Tasgius" box="[532,757,997,1019]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="B0F3636EF906FFC1FDC5FC23FD4DFC7B" box="[532,649,997,1019]" italics="true" pageId="23" pageNumber="224">Staphylinus</emphasis>
(
<emphasis id="B0F3636EF906FFC1FD4EFC20FD2FFC7B" box="[671,747,998,1019]" italics="true" pageId="23" pageNumber="224">Tasgius</emphasis>
)
</taxonomicName>
; description); 
<bibRefCitation id="E616C28DF906FFC1FC44FC23FB90FC7B" box="[917,1108,997,1019]" pageId="23" pageNumber="224" refId="ref33499" refString="Scheerpeltz, O. (1933) Staphylinidae VII (Pars 129). Supplementum I. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum Catalogus. Vol. VI. Staphylinidae. Junk, Berlin, pp. 989 - 1500." type="book chapter">Scheerpeltz, 1933</bibRefCitation>
:1391 (
<taxonomicName id="4587C4FFF906FFC1FB73FC23FAD3FC7B" box="[1186,1303,997,1019]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="B0F3636EF906FFC1FB73FC23FAD3FC7B" box="[1186,1303,997,1019]" italics="true" pageId="23" pageNumber="224">Staphylinus</emphasis>
</taxonomicName>
; catalogue);
</paragraph>
<paragraph id="8238BF7CF906FFC1FF16FBC2FA58FBB8" blockId="23.[151,1437,752,1142]" pageId="23" pageNumber="224">
<bibRefCitation id="E616C28DF906FFC1FF16FBC2FE0CFB9A" box="[199,456,1028,1050]" pageId="23" pageNumber="224" refId="ref34111" refString="Smetana, A. &amp; Davies A. (2000) Reclassification of the north temperate taxa associated with Staphylinus sensu lato, including comments on relevant subtribes of Staphylinini (Coleoptera: Staphylinidae). American Museum Novitates, 3287, 1 - 88. https: // doi. org / 10.1206 / 0003 - 0082 (2000) 287 % 3 C 0001: ROTNTT % 3 E 2.0. CO; 2" type="journal article">Smetana &amp; Davies, 2000</bibRefCitation>
: 33 (
<taxonomicName id="4587C4FFF906FFC1FE28FBC2FD81FB99" authorityName="Smetana" authorityYear="2018" box="[505,581,1028,1049]" class="Insecta" family="Staphylinidae" genus="Ocypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="B0F3636EF906FFC1FE28FBC2FD81FB99" box="[505,581,1028,1049]" italics="true" pageId="23" pageNumber="224">Ocypus</emphasis>
</taxonomicName>
; homonym) 
<emphasis id="B0F3636EF906FFC1FD19FBC2FCFFFB9A" bold="true" box="[712,827,1028,1050]" pageId="23" pageNumber="224">comb. nov.</emphasis>
, 
<emphasis id="B0F3636EF906FFC1FC98FBC2FC62FB9A" bold="true" box="[841,934,1028,1050]" pageId="23" pageNumber="224">syn. nov.</emphasis>
<taxonomicName id="4587C4FFF906FFC1FF46FBE5FDE6FBB8" authority="Smetana &amp; Davies, 2000: 33" authorityName="Smetana &amp; DaVies" authorityPageNumber="33" authorityYear="2000" box="[151,546,1058,1080]" class="Insecta" family="Staphylinidae" genus="Ocypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="cameroni">
<emphasis id="B0F3636EF906FFC1FF46FBE5FF32FBB8" box="[151,246,1059,1080]" italics="true" pageId="23" pageNumber="224">cameroni</emphasis>
<bibRefCitation id="E616C28DF906FFC1FF2DFBE4FE3BFBB8" box="[252,511,1058,1080]" pageId="23" pageNumber="224" refId="ref34111" refString="Smetana, A. &amp; Davies A. (2000) Reclassification of the north temperate taxa associated with Staphylinus sensu lato, including comments on relevant subtribes of Staphylinini (Coleoptera: Staphylinidae). American Museum Novitates, 3287, 1 - 88. https: // doi. org / 10.1206 / 0003 - 0082 (2000) 287 % 3 C 0001: ROTNTT % 3 E 2.0. CO; 2" type="journal article">Smetana &amp; Davies, 2000</bibRefCitation>
: 33
</taxonomicName>
(
<taxonomicName id="4587C4FFF906FFC1FDE3FBE5FDBAFBB8" authorityName="Smetana" authorityYear="2018" box="[562,638,1059,1080]" class="Insecta" family="Staphylinidae" genus="Ocypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="B0F3636EF906FFC1FDE3FBE5FDBAFBB8" box="[562,638,1059,1080]" italics="true" pageId="23" pageNumber="224">Ocypus</emphasis>
</taxonomicName>
; replacement name for 
<taxonomicName id="4587C4FFF906FFC1FCA1FBE4FACAFBB8" authority="Cameron, 1932" authorityName="Cameron" authorityYear="1932" box="[880,1294,1058,1080]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="bimaculatus">
<emphasis id="B0F3636EF906FFC1FCA1FBE4FBACFBB8" box="[880,1128,1058,1080]" italics="true" pageId="23" pageNumber="224">Staphylinus bimaculatus</emphasis>
Cameron, 1932
</taxonomicName>
, nec Schrank,
</paragraph>
<paragraph id="8238BF7CF906FFC1FF16FB87FA59FBD7" blockId="23.[151,1437,752,1142]" box="[199,1437,1089,1111]" pageId="23" pageNumber="224">
1798); 
<bibRefCitation id="E616C28DF906FFC1FEC2FB84FE6CFBD7" box="[275,424,1089,1111]" pageId="23" pageNumber="224" refId="ref32612" refString="Herman, L. H. (2001) Catalogue of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. VI. Staphylinine group. Part 3. Staphylininae: Staphylinini (Quediina, Staphylinina, Tanygnathinina, Xanthopygina). Xantholinini. Staphylinidae incertae sedis, fossils. Protactinae. Bulletin of the American Museum of Natural History, 265, 3021 - 3840." type="journal article">Herman, 2001</bibRefCitation>
:3370 (
<taxonomicName id="4587C4FFF906FFC1FE23FB84FDFAFBD7" authorityName="Smetana" authorityYear="2018" box="[498,574,1090,1111]" class="Insecta" family="Staphylinidae" genus="Ocypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="B0F3636EF906FFC1FE23FB84FDFAFBD7" box="[498,574,1090,1111]" italics="true" pageId="23" pageNumber="224">Ocypus</emphasis>
</taxonomicName>
; catalogue); 
<bibRefCitation id="E616C28DF906FFC1FD18FB87FCA3FBD7" box="[713,871,1089,1111]" pageId="23" pageNumber="224" refId="ref34017" refString="Smetana, A, (2004) Staphylininae. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Apollo Books, Stenstrup, pp. 624 - 698." type="book chapter">Smetana, 2004</bibRefCitation>
: 676 (
<taxonomicName id="4587C4FFF906FFC1FC7DFB84FC3CFBD7" authorityName="Smetana" authorityYear="2018" box="[940,1016,1090,1111]" class="Insecta" family="Staphylinidae" genus="Ocypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="B0F3636EF906FFC1FC7DFB84FC3CFBD7" box="[940,1016,1090,1111]" italics="true" pageId="23" pageNumber="224">Ocypus</emphasis>
</taxonomicName>
; catalogue); 
<bibRefCitation id="E616C28DF906FFC1FB52FB87FA5CFBD7" box="[1155,1432,1089,1111]" pageId="23" pageNumber="224" refId="ref33592" refString="Schulke, M. &amp; Smetana, A. (2015) Staphylinina. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Revised and updated Edition. Brill, Leiden, pp. 1081 - 1107." type="book chapter">Schülke &amp; Smetana, 2015</bibRefCitation>
:
</paragraph>
<paragraph id="8238BF7CF906FFC1FF16FBA6FD6AFBF6" blockId="23.[151,1437,752,1142]" box="[199,686,1120,1142]" pageId="23" pageNumber="224">
1089 (
<taxonomicName id="4587C4FFF906FFC1FED9FBA7FE90FBF6" authorityName="Smetana" authorityYear="2018" box="[264,340,1121,1142]" class="Insecta" family="Staphylinidae" genus="Ocypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="B0F3636EF906FFC1FED9FBA7FE90FBF6" box="[264,340,1121,1142]" italics="true" pageId="23" pageNumber="224">Ocypus</emphasis>
</taxonomicName>
, catalogue
<emphasis id="B0F3636EF906FFC1FE13FBA6FDF9FBF6" bold="true" box="[450,573,1120,1142]" pageId="23" pageNumber="224">) comb. nov</emphasis>
., 
<emphasis id="B0F3636EF906FFC1FD80FBA6FD6AFBF6" bold="true" box="[593,686,1120,1142]" pageId="23" pageNumber="224">syn. nov.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="CA9DECF7F906FFC1FF46FB59FC21FA55" pageId="23" pageNumber="224" type="materials_examined">
<paragraph id="8238BF7CF906FFC1FF46FB59FC46FB38" blockId="23.[151,1437,1183,1494]" box="[151,898,1183,1208]" pageId="23" pageNumber="224">
<emphasis id="B0F3636EF906FFC1FF46FB59FE81FB38" bold="true" box="[151,325,1183,1208]" pageId="23" pageNumber="224">
<typeStatus id="5D3C01DEF906FFC1FF46FB59FF16FB38" box="[151,210,1183,1208]" pageId="23" pageNumber="224">Type</typeStatus>
material.
</emphasis>
<taxonomicName id="4587C4FFF906FFC1FE9DFB59FC46FB38" authority="Bernhauer, 1920" authorityName="Bernhauer" authorityYear="1920" box="[332,898,1183,1208]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="sikkimensis" subGenus="Tasgius">
<emphasis id="B0F3636EF906FFC1FE9DFB59FE0AFB38" box="[332,462,1183,1208]" italics="true" pageId="23" pageNumber="224">Staphylinus</emphasis>
(
<emphasis id="B0F3636EF906FFC1FE0CFB67FDEBFB38" box="[477,559,1185,1208]" italics="true" pageId="23" pageNumber="224">Tasgius</emphasis>
) 
<emphasis id="B0F3636EF906FFC1FD93FB59FD06FB38" box="[578,706,1183,1208]" italics="true" pageId="23" pageNumber="224">sikkimensis</emphasis>
<bibRefCitation id="E616C28DF906FFC1FD18FB59FC46FB38" box="[713,898,1183,1208]" pageId="23" pageNumber="224" refId="ref31990" refString="Bernhauer, M. (1920) Neue Staphyliniden des indo-malaischen Gebietes. Archiv fur Naturgeschichte (A), 84, 177 - 188." type="journal article">Bernhauer, 1920</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8238BF7CF906FFC1FF16FB07FD55FB5A" blockId="23.[151,1437,1183,1494]" box="[199,657,1217,1242]" pageId="23" pageNumber="224">
<materialsCitation id="32EFB521F906FFC1FF16FB07FD55FB5A" ID-GBIF-Occurrence="1913888599" box="[199,657,1217,1242]" country="India" location="Sikkim" pageId="23" pageNumber="224" specimenCount="1" stateProvince="Sikkim" typeStatus="holotype">
<emphasis id="B0F3636EF906FFC1FF16FB07FEA1FB5A" bold="true" box="[199,357,1217,1242]" pageId="23" pageNumber="224">
<typeStatus id="5D3C01DEF906FFC1FF16FB07FEC6FB5A" box="[199,258,1217,1242]" pageId="23" pageNumber="224">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="FA90FFECF906FFC1FEBDFB07FE6EFB5A" box="[364,426,1217,1242]" name="India" pageId="23" pageNumber="224">India</collectingCountry>
: 
<collectingRegion id="4043719EF906FFC1FE65FB07FDCCFB5A" box="[436,520,1217,1242]" country="India" name="Sikkim" pageId="23" pageNumber="224">Sikkim</collectingRegion>
, Darjeeling.
</materialsCitation>
</paragraph>
<paragraph id="8238BF7CF906FFC1FF16FB23FC21FA55" blockId="23.[151,1437,1183,1494]" pageId="23" pageNumber="224">
<emphasis id="B0F3636EF906FFC1FF16FB23FE18FB7E" bold="true" box="[199,476,1253,1278]" pageId="23" pageNumber="224">Type specimen studied</emphasis>
. 
<emphasis id="B0F3636EF906FFC1FE3CFB23FD9EFB7E" bold="true" box="[493,602,1253,1278]" pageId="23" pageNumber="224">Holotype</emphasis>
(male) by mοnοtypy, in FMNH: “Sikkim / Darjeeling / Christie / n.sp. Désiré me manqué (handwritten) / Ich habe nicht (bei aureοguttatus Fvl. Ind. (handwritten) / Bestimmung vοn Fauvel (handwritten in red ink) / 
<taxonomicName id="4587C4FFF906FFC1FDE2FAEBFCCBFAC6" authority="Bernh." authorityName="Bernhauer" authorityYear="1920" box="[563,783,1325,1350]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="sikkimensis">sikkimensis Bernh.</taxonomicName>
Typus un. Wasmanni / Chicagο NHMus M. Bernhauer Cοllectiοn / Hοlοtypus ♂ 
<taxonomicName id="4587C4FFF906FFC1FE1BFA97FCFCFAE9" authority="Bh. K." authorityName="Bh. K." box="[458,824,1361,1386]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="sikkimensis">Staphylinus sikkimensis Bh. K.</taxonomicName>
Grebennikοv det. 2001 (handwritten οn red label) / 
<taxonomicName id="4587C4FFF906FFC1FF46FAB0FF28FA0E" authority="Smetana, 2018" authorityName="Smetana" authorityYear="2018" box="[151,236,1398,1422]" class="Insecta" family="Staphylinidae" genus="Ocypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus" sensu="stricto">Ocypus</taxonomicName>
<taxonomicNameLabel id="ABC0DE15F906FFC1FF25FAB0FEECFA0D" box="[244,296,1398,1421]" pageId="23" pageNumber="224" sensu="stricto">s.str.</taxonomicNameLabel>
♂ 
<taxonomicName id="4587C4FFF906FFC1FE9DFAB3FDD6FA0E" authority="(Bh.)" authorityName="Bh." box="[332,530,1397,1422]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="sikkimensis">sikkimensis (Bh.)</taxonomicName>
K. A. Grebennikοv det. 2001 (handwritten οn white label). The genital segment and the aedοeagus are οn twο separate transparent plates attached tο the specimen. The specimen is in fair shape, οnly three segments οf the right antenna and twο οf the left οne are present.
</paragraph>
</subSubSection>
</treatment>
</document>