Systematic revision of American Glaresidae (Coleoptera: Scarabaeoidea) Author Gordon, Robert D. Author Hanley, Guy A. text Insecta Mundi 2014 2014-01-10 2014 333 1 91 journal article 10.5281/zenodo.4645613 1942-1354 4645613 112A1F0B-1A82-4672-842B-A79A21F251D6 Glaresis costata Gordon and Hanley , new species Description . Male . Length 3.7 mm , width 1.9 mm ; body form elongate, slightly widened from elytral base to apical 1/3 ( Fig. 4A ). Color yellowish brown. Head with clypeus and frons densely microreticulate, dull, with large, deep punctures, setae short, barely emergent from punctures; frons coarsely, densely microreticulate, feebly shiny, with few small, round tubercles, not setose; vertex with short, median basal carina, surface coarsely, densely microreticulate, feebly shiny, without tubercles or punctures. Clypeal apex truncate, smooth, narrow, angled posteriorly, lateral angles abruptly angulate ( Fig. 4B ). Mandible pair symmetrical; mesal tooth strong; lateral prominence strong, pronounced; outer margin angular. Pronotum with distinct central furrow extended from base nearly to apex, surface dull, coarsely, densely micoreticulate, with elongate, setae-bearing ridges arranged in various directions, setae distinct, slightly shorter than ridge ( Fig. 4C ). Anterior and lateral pronotal margins not bordered, anterior margin smooth medially, crenulate in lateral 1/4, lateral and basal margins entirely crenulate, basal margin bordered. Elytron with convex, somewhat carinate striae, surface feebly shiny, coarsely, densely microreticulate; each stria with small, closely spaced, nearly round, seta-bearing tubercle (presenting slightly carinate appearance), setae long, slightly longer than diameter of an interval puncture; interval with large, distinctly visible, rectangular punctures ( Fig. 4E ). Metasternum long, feebly shiny, coarsely, densely microreticulate throughout; entire surface with short, oblique, seta-bearing ridges; without trace of metasternal groove. Lateral protibial teeth unevenly spaced, basal tooth reduced. Mesotibia with 5 spines laterally, spinal bases widely spaced, outer apex of tibia not strongly projecting ( Fig. 4D ). Posterior margin of metatrochanter with 2 large teeth (usually an additional small tooth present) ( Fig. 4H ); posterosuperior surface without teeth. Metafemoral surface with widely scattered, elongate, setae-bearing ridges, finely microreticulate, dull throughout; shape slightly elongate; width to length ratio1.0:1.6, without flange on anterior margin; posterosuperior margin with single tooth ( Fig. 4H ). Metatibial surface finely microreticulate except extreme apex, and apex of lateral expansion shiny (probably from wear). Outer lateral metatibial margin serrate with row of small, setae-bearing teeth, posteromedian lateral projection large, shelf-like, apical margin uneven; row of more or less 3 tubercles anterior to lateral expansion; median surface and apex without tubercles; inner lateral margin with row of small, setabearing tubercles, emarginate in apical 1/3, small tooth present at base of emargination ( Fig. 4G ). Apex of 5th abdominal ventrite truncate. Genitalia long, dorso-ventrally flattened; basal piece shorter than parameres, proximal end curved; parameres straight in lateral view; median lobe longer than parameres, slightly wider at middle than a paramere, strongly curved upward in apical 1/3, apex rounded in ventral and lateral views ( Fig. 4F ). Female . Apex of 5th abdominal ventrite same as in male. Variation . Length 3.0 to 3.7 mm , width 1.7 to 2.0 mm. Carina on vertex may be absent; pronotal ridges vary somewhat in degree of spacing; surface sculpture varies slightly, clypeus and anterior portion of frons often shiny, probably because of wear; posterior trochanteral margin with 2 or 3 teeth, inner most tooth small, often not detectable. Type material . Holotype male; Colorado : Hasty ( Bent County ), Colorado , 14.VI..82, Lot 11, BG& JL Carr ( JCIC ) . Paratypes , 113: (46) same data as holotype except date 9.VI.83. Colorado : (1) Nunn , Pawnee Grassland , 6-VI-1977 , J.W. Leetham ; (24) New Mexico : Chaves Co. , 35.5 mi. E. Roswell , VII-31- 1978 ; (2) Otero Co.; Three Rivers Petroglyphs , 7-VI-1987 , Robert Gordon. Texas : (34) Hudspeth Co. , 2mi N Fort Hancock , V-29-1982 , V-30-1982 , N. Rulien ; (4) 10 mi. N. Andrews ( Andrews Co. ), 13.VI.83, Lot 2, BF& JL Carr ; (1) Monahans ( Ward Co. ), 20.IV.87, BF& JL Carr. ( CNIC ) ( JCIC ) ( FSCA ) ( MJPC ) ( USNM ) . Other specimens . 18: (1) New Mexico : Eddy Co. , 32 o 24.8’N , 103 o 41.5’W ; Eddy Co. , 26 mi. N Carlsbad. (1) South Dakota : Angostura Dam, S. of Hot Springs , Vii 6-10, 1968 , H.F. Howden. (1) Texas : El Paso Co. , 12 mi. NE Fabans , IV-23-1996 , UV light, Coll. E. G. Riley-628; (2) Utah : (1) UTAH , Emery Co. , Goblin Valley Rd .; Utah , Emery Co. 5100', 4airMiN GilsonBt , VII-20/23-81, Viers , Parker , Griswold ; Emery Co. , Goblin Valley , 4-VIII-1986 , Robert Gordon ; (1) Utah , Emery Co. , Little Gilson Butte , 29-VIII-1986 , A. S. Menke ; (4) Utah , Hanksville , VIII 7-9, 1968 , H.F. Howden ; (1) Utah , 14 mi. S. Hanksville , 25-VII-1968 , J.E. Martin ; (1) Fairview Ranch , 5000', 13 mi. S. Hanksville , 6.VIII.1968 ; (1) Utah , 26 mi. S. Hanksville ( Garfield Co. ), Hwy. 95, 16-VI-84, Robert Gordon ; Grand Co. , 3 mi north of Moab , Hwy 123. ( BYUC ) ( CMNC ) ( CNIC ) ( MJPC ) ( TAMU ) ( USNM ) ( USUL ) . Remarks . In addition to key characters, G . costata differs from G . ecostata by the more roughly microsculptured body surface, distinct median pronotal furrow, bordered basal pronotal margin, and long metasternal setae. Etymology . The name ecostata refers to a lack of elytral costae. Therefore, this species is named G . costata in reference to the slightly carinate appearance of the elytral striae, and in recognition of its close relationship to G . ecostata .