Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda)
Author
Korshunova, Tatiana
Author
Martynov, Alexander
Author
Bakken, Torkild
Author
Evertsen, Jussi
Author
Fletcher, Karin
Author
Mudianta, I Wayan
Author
Saito, Hiroshi
Author
Lundin, Kennet
Author
Michael Schroedl,
Author
Picton, Bernard
text
ZooKeys
2017
717
1
139
http://dx.doi.org/10.3897/zookeys.717.21885
journal article
http://dx.doi.org/10.3897/zookeys.717.21885
1313-2970-717-1
C19B43B1B3214CB1B1B2A246CEAC56BC
Calmella
Eliot, 1910
Figs 33, 34
Type
species.
Eolidia cavolini
Verany
, 1846.
Diagnosis.
Body narrow. Notal edge completely absent. Cerata on short stalks in several groups. Rhinophores smooth, larger than oral tentacles or similar in size. Anterior foot corners present. Anus mixed: pleuroproctic shifted towards dorsal acleioproctic position. Distinct oral glands. Rachidian teeth with narrow compressed cusp and distinct denticles. Lateral teeth weakly denticulated to smooth with attenuated process basally. Single distal receptaculum seminis. Moderately long prostatic non-granulated vas deferens. Penis conical.
Species included.
Calmella cavolini
(
Verany
, 1846) (Fig. 33) (original description in
Verany
1846, detailed description in
Schmekel and Portmann 1982
),
Calmella gaditana
(Cervera,
Garcia-Gomez
&
Garcia
, 1987), comb. n. (original description in
Cervera et al. 1987
, = "
Flabellina
" confusa
Gonzalez-Duarte
et al. 2008
, original description in
Gonzalez-Duarte
et al. 2008
; for discussion see
Furfaro et al. 2017
),
Calmella bandeli
Marcus, 1976 (original description in
Marcus 1976
).
Remarks.
The genus
Calmella
was established by
Eliot (1910)
and since that time it has never been synonymised with
Flabellina
, until the most recent
Furfaro et al. (2017)
study. According to the present phylogenetic analysis
Calmella
is valid, related to the true
Flabellina
s. str. (Fig. 2) and differs morphologically regarding its much less ramified and raised ceratal stalks and presence of smooth rhinophores. See remarks also under the genus
Piseinotecus
.