Middle Ordovician (Darriwilian) cheirurid trilobites from the Table Cove Formation, western Newfoundland, Canada
Author
Adrain, Jonathan M.
0000-0002-7000-1311
jonathan-adrain@uiowa.edu
Author
Pérez-Peris, Francesc
0000-0002-7000-1311
jonathan-adrain@uiowa.edu
text
Zootaxa
2021
2021-09-24
5041
1
1
73
http://dx.doi.org/10.11646/zootaxa.5041.1.1
journal article
10.11646/zootaxa.5041.1.1
1175-5326
5531880
5E82BE60-609F-4287-AC67-D86536FB7686
Kawina
Barton, 1915
=
Kolymella
Čugaeva, 1973
Type
species.
By
original designation;
Cheirurus vulcanus
Billings, 1865a
, from the
Shallow Bay
Formation
(
lower Darriwilian
) of
western Newfoundland
,
Canada
(
Laurentia
)
.
Other species.
Kawina arnoldi
Whittington, 1963
, Shallow Bay Formation (lower Darriwilian), Newfoundland,
Canada
(Laurentia);
K. limbata
Whittington, 1963
, Shallow Bay Formation (lower Darriwilian), Newfoundland,
Canada
(Laurentia);
K. plana
Čugaeva
in
Čugaeva
et al
., 1964
, Elgenchak Formation (Darriwilian),
Magadan Oblast
, far eastern
Russia
(Kolyma-Omolon Terrane);
K. spinigena
Ancygin, 1977
, Karakol-Mikhailovsk Horizon (Darriwilian), Urals,
Russia
(Baltica);
K. stougei
n. sp.
, Table Cove Formation (Darriwilian), Newfoundland,
Canada
(Laurentia);
K. wilsoni
Ross, 1972
, Antelope Valley Formation (Dapingian),
Nevada
,
USA
(Laurentia).
Diagnosis.
Glabella widest posteriorly, across middle or anterior portion of L1; librigenal lateral border strongly inflated; librigenal lateral border furrow narrow and field broad; hypostome with long anterior lobe of middle body, middle furrows weakly expressed and set far posteriorly; pygidium with three well expressed axial rings, axial furrows meeting posteriorly to fully circumscribe axis, and broad, dorsoventrally flattened, pleural spines with blunt ends; pleural spines attached to adjacent spines for much of their length.
Discussion.
Restriction of
Kawina
to a core Middle Ordovician group sharing all of the prominent diagnostic features and transfer of some previously assigned species with question to
Cydonocephalus
were discussed above. Most of the species are Laurentian, but the Darriwilian
K. spinigena
Ancygin, 1977
, is from Baltica and shares all of the cranidial and pygidial synapomorphies of the genus.
Kawina plana
Čugaeva
in
Čugaeva
et al
., 1964
,
type
species of the monotypic
Kolymella
Čugaeva, 1973
, also shares the cranidial and pygidial characters of
Kawina
. This species is from the Elgenchak Formation (Darriwilian), Elgenchak Mountains,
Magadan Oblast
, far eastern
Russia
. The faunas of the Kolyma-Omolon Terrane contain a mix of endemics, taxa with Siberian affinity, and taxa with Laurentian affinity (see discussion of
Parasphaerexochus
above).
Kolymella
is one of the latter, as the only feature that seems to differentiate it from species of
Kawina
is the apparent lack of inflation of the glabella (
Fortey, 1980
, p. 86). Exactly how much the glabella is dorsally inflated however, is difficult to judge, as Čugaeva (
Čugaeva
et al
.,1964
;
Čugaeva, 1973
) did not illustrate any lateral or anterior views. Nevertheless, one slightly oblique photograph (
Čugaeva, 1973
, pl. 10, fig. 2b) does suggest fairly low dorsal convexity. Given that this is the only point of differentiation, in the current state of knowledge we regard it as a specific autapomorphy of
K. plana
and consider
Kolymella
a junior subjective synonym of
Kawina
.