<document id="FF8A63A7C2E43DEDEBF30EB3E85CA759" ID-DOI="10.5852/ejt.2020.718.1101" ID-Zenodo-Dep="4069574" ID-ZooBank="F9E9A91E-488C-4DB1-9361-E788E9AC5BC1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1602021550335" checkinUser="valdenar" docAuthor="Huber, Bernhard A. &amp; Villarreal, Osvaldo" docDate="2020" docId="03E887ADFEC87B19FD1FFE44F9C3FCFA" docLanguage="en" docName="ejt-718_huber_villareal.pdf.imf" docOrigin="European Journal of Taxonomy 718" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Systenita Simon 1893" docType="treatment" docVersion="7" lastPageNumber="281" masterDocId="FFD1FFD5FFD07A01FF8BFF8FFA5BFFBA" masterDocTitle="On Venezuelan pholcid spiders (Araneae, Pholcidae)" masterLastPageNumber="317" masterPageNumber="1" pageNumber="281" updateTime="1698792674692" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="1B97191DD27649387B85047491228DE3">On Venezuelan pholcid spiders (Araneae, Pholcidae)</mods:title>
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<mods:namePart id="52DD0428A1CAF488932BB1FDDB17C710">Huber, Bernhard A.</mods:namePart>
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<mods:affiliation id="C60A10B4436FA61209FBEFFA54D82AD7">Zoological Research Museum Alexander Koenig, Bonn, Germany.</mods:affiliation>
<mods:nameIdentifier id="0C3DAC7BA478CEFB60FA793FAB8CF26B" type="email">b.huber@leibniz-zfmk.de</mods:nameIdentifier>
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<mods:namePart id="18565146127DD1F1EE92D6BC440061DF">Villarreal, Osvaldo</mods:namePart>
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<mods:affiliation id="083EE52D07A505DB8B58256E379827C0">Museo del Instituto de Zoología Agrícola, Universidad Central de Venezuela, Maracay, Venezuela. &amp; Museu Nacional / UFRJ, Rio de Janeiro, Brazil.</mods:affiliation>
<mods:nameIdentifier id="C0FD26532E3B7C5A434972E588C8AAC7" type="email">osvaldovillarreal@gmail.com</mods:nameIdentifier>
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<mods:title id="94464574FC54F77EA22FE1CE38AA7E92">European Journal of Taxonomy</mods:title>
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<mods:date id="C1CF3970F948EBCFED2972A6C4E591B7">2020</mods:date>
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<paragraph id="8BFE36BBFEC87B19FD1FFE44F9C4FE5F" blockId="280.[660,927,459,485]" box="[660,927,459,485]" pageId="280" pageNumber="281">
<heading id="D0B681D7FEC87B19FD1FFE44F9C4FE5F" box="[660,927,459,485]" centered="true" fontSize="11" level="2" pageId="280" pageNumber="281" reason="2">
<taxonomicName id="4C414D38FEC87B19FD1FFE44F9C4FE5F" ID-CoL="7RN7" ID-ENA="280444" authority="Simon, 1893" authorityName="Simon" authorityYear="1893" box="[660,927,459,485]" class="Arachnida" family="Pholcidae" genus="Systenita" kingdom="Animalia" order="Araneae" pageId="280" pageNumber="281" phylum="Arthropoda" rank="genus">
<emphasis id="B935EAA9FEC87B19FD1FFE44F95AFE5F" bold="true" box="[660,769,459,485]" italics="true" pageId="280" pageNumber="281">Systenita</emphasis>
Simon, 1893
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</paragraph>
<paragraph id="8BFE36BBFEC87B19FF36FD80FB58FD93" blockId="280.[189,259,527,553]" box="[189,259,527,553]" pageId="280" pageNumber="281">
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<emphasis id="B935EAA9FEC87B19FF36FD80FB58FD93" bold="true" box="[189,259,527,553]" pageId="280" pageNumber="281">Notes</emphasis>
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<paragraph id="8BFE36BBFEC87B19FF36FDCEF9C3FCFA" blockId="280.[189,1399,576,832]" pageId="280" pageNumber="281">
<taxonomicName id="4C414D38FEC87B19FF36FDCEFB7CFDE1" authorityName="Simon" authorityYear="1893" box="[189,295,577,603]" class="Arachnida" family="Pholcidae" genus="Systenita" kingdom="Animalia" order="Araneae" pageId="280" pageNumber="281" phylum="Arthropoda" rank="genus">
<emphasis id="B935EAA9FEC87B19FF36FDCEFB7CFDE1" box="[189,295,577,603]" italics="true" pageId="280" pageNumber="281">Systenita</emphasis>
</taxonomicName>
continues to be a monotypic genus. The newly described 
<taxonomicName id="4C414D38FEC87B19FC5DFDCFFF48FDE0" authority="Huber" authorityName="Huber" authorityYear="2020" box="[982,1299,576,602]" class="Arachnida" family="Pholcidae" genus="Mecolaesthus" kingdom="Animalia" order="Araneae" pageId="280" pageNumber="281" phylum="Arthropoda" rank="species" species="fallax" status="sp. nov.">
<emphasis id="B935EAA9FEC87B19FC5DFDCFFE99FDE0" box="[982,1218,576,602]" italics="true" pageId="280" pageNumber="281">Mecolaesthus fallax</emphasis>
Huber
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<taxonomicNameLabel id="A20657D2FEC87B19FA97FDCEFF2CFDE1" box="[1308,1399,577,603]" pageId="280" pageNumber="281" rank="species">sp. nov.</taxonomicNameLabel>
is superficially extremely similar (indistinguishable in the field) but fundamentally different in many details of genital morphology. Our preliminary molecular data (J.J. Astrin, B.A. Huber, unpubl.data) place 
<taxonomicName id="4C414D38FEC87B19FE89FD3CFB97FD77" authorityName="Simon" authorityYear="1893" box="[258,460,691,717]" class="Arachnida" family="Pholcidae" genus="Systenita" kingdom="Animalia" order="Araneae" pageId="280" pageNumber="281" phylum="Arthropoda" rank="species" species="prasina">
<emphasis id="B935EAA9FEC87B19FE89FD3CFB97FD77" box="[258,460,691,717]" italics="true" pageId="280" pageNumber="281">Systenita prasina</emphasis>
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closer to the new genus 
<taxonomicName id="4C414D38FEC87B19FD61FD3BF9FFFD77" authority="Huber" authorityName="Huber &amp; Villarreal" authorityYear="2020" box="[746,932,691,717]" class="Arachnida" family="Pholcidae" genus="Boconita" kingdom="Animalia" order="Araneae" pageId="280" pageNumber="281" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B935EAA9FEC87B19FD61FD3BF90FFD77" box="[746,852,692,717]" italics="true" pageId="280" pageNumber="281">Boconita</emphasis>
Huber
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<taxonomicNameLabel id="A20657D2FEC87B19FC20FD3CFE4EFD77" box="[939,1045,691,717]" pageId="280" pageNumber="281" rank="genus">gen. nov.</taxonomicNameLabel>
and to 
<taxonomicName id="4C414D38FEC87B19FBE1FD3CFAA3FD49" authority="Simon, 1893" authorityName="Simon" authorityYear="1893" class="Arachnida" family="Pholcidae" genus="Coryssocnemis" kingdom="Animalia" order="Araneae" pageId="280" pageNumber="281" phylum="Arthropoda" rank="genus">
<emphasis id="B935EAA9FEC87B19FBE1FD3CFF47FD77" box="[1130,1308,691,717]" italics="true" pageId="280" pageNumber="281">Coryssocnemis</emphasis>
Simon, 1893
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than to 
<taxonomicName id="4C414D38FEC87B19FEEBFD56F8FAFD49" authority="Huber" authorityName="Huber" authorityYear="2020" box="[352,673,729,755]" class="Arachnida" family="Pholcidae" genus="Mecolaesthus" kingdom="Animalia" order="Araneae" pageId="280" pageNumber="281" phylum="Arthropoda" rank="species" species="fallax" status="sp. nov.">
<emphasis id="B935EAA9FEC87B19FEEBFD56F815FD49" box="[352,590,729,755]" italics="true" pageId="280" pageNumber="281">Mecolaesthus fallax</emphasis>
Huber
</taxonomicName>
<taxonomicNameLabel id="A20657D2FEC87B19FD27FD55F952FD4E" box="[684,777,730,756]" pageId="280" pageNumber="281" rank="species">sp. nov.</taxonomicNameLabel>
The superficial similarity with 
<taxonomicName id="4C414D38FEC87B19FB03FD56FB5DFCA3" authority="Huber" authorityName="Huber" authorityYear="2020" class="Arachnida" family="Pholcidae" genus="Mecolaesthus" kingdom="Animalia" order="Araneae" pageId="280" pageNumber="281" phylum="Arthropoda" rank="species" species="fallax" status="sp. nov.">
<emphasis id="B935EAA9FEC87B19FB03FD56FF2CFD49" box="[1160,1399,729,755]" italics="true" pageId="280" pageNumber="281">Mecolaesthus fallax</emphasis>
Huber
</taxonomicName>
<taxonomicNameLabel id="A20657D2FEC87B19FE84FC8FFB30FCA0" box="[271,363,768,794]" pageId="280" pageNumber="281" rank="species">sp. nov.</taxonomicNameLabel>
is thus most probably a result of convergent evolution due to microhabitat change and adaptation to the same microhabitat (underside of live leaves).
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