<document id="BFD53DFE6BE9A5EC4D99A333896031BA" ID-CLB-Dataset="298900" ID-DOI="10.1643/ci-18-055" ID-GBIF-Dataset="4919bfd6-3d35-48b9-9421-c852fa35abab" ID-Zenodo-Dep="12536069" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.metadata_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1719340789171" checkinUser="julia" docAuthor="Ma, Ka Yan &amp; Craig, Matthew T." docDate="2018" docId="03FEC024E12BDF3EFCD738BE6954FBC7" docLanguage="en" docName="copeia.106.3.443-456.pdf" docOrigin="Copeia 106 (3)" docSource="http://dx.doi.org/10.1643/ci-18-055" docTitle="Variola Swainson 1839" docType="treatment" docVersion="5" lastPageNumber="456" masterDocId="FFC7B85CE126DF33FFE33B236D19FFA4" masterDocTitle="An Inconvenient Monophyly: An Update on the Taxonomy of the Groupers (Epinephelidae)" masterLastPageNumber="456" masterPageNumber="443" pageNumber="456" updateTime="1719854615312" updateUser="julia">
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<mods:title id="4D0828DCDA240BFC402E9930F98E8F53">An Inconvenient Monophyly: An Update on the Taxonomy of the Groupers (Epinephelidae)</mods:title>
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<mods:namePart id="B7AD8E598515E59CAC19C38E39330FEA">Ma, Ka Yan</mods:namePart>
<mods:affiliation id="9BE7803EB41A0A4BA82C7DA5730618F2">Simon F. S. Li Marine Science Laboratory, School of Life Sciences, The Chinese University of Hong Kong, Sha Tin, 999077, Hong Kong SAR.</mods:affiliation>
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<mods:namePart id="751C847135A1F6DA721E0C5AD296FA71">Craig, Matthew T.</mods:namePart>
<mods:affiliation id="EB44CD2EF203E37B9D936895A44B437C">2 National Oceanic and Atmospheric Administration, National Marine Fisheries Service, Southwest Fisheries Science Center, 8901 La Jolla Shores Drive, La Jolla, California 92037</mods:affiliation>
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<mods:date id="857CBD6D299C109B66913C3BD3A148E0">2018</mods:date>
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<treatment id="03FEC024E12BDF3EFCD738BE6954FBC7" ID-DOI="http://doi.org/10.5281/zenodo.12536351" ID-GBIF-Taxon="233767938" ID-Zenodo-Dep="12536351" LSID="urn:lsid:plazi:treatment:03FEC024E12BDF3EFCD738BE6954FBC7" httpUri="http://treatment.plazi.org/id/03FEC024E12BDF3EFCD738BE6954FBC7" lastPageNumber="456" pageId="13" pageNumber="456">
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<paragraph id="8BE87132E12BDF3EFCD738BE69DBFC10" blockId="13.[820,1490,925,1123]" box="[820,1218,925,948]" pageId="13" pageNumber="456">
<heading id="D0A0C65EE12BDF3EFCD738BE69DBFC10" bold="true" box="[820,1218,925,948]" fontSize="9" level="3" pageId="13" pageNumber="456" reason="6">
<emphasis id="B923AD20E12BDF3EFCD738BE69DBFC10" bold="true" box="[820,1218,925,948]" pageId="13" pageNumber="456">
Genus 
<taxonomicName id="4C570AB1E12BDF3EFC6038BE6958FC10" ID-CoL="85YG" authority="Swainson" authorityName="Swainson" authorityYear="1839" box="[899,1089,925,948]" family="Serranidae" genus="Variola" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="456" phylum="Chordata" rank="genus">
<emphasis id="B923AD20E12BDF3EFC6038BE6EC9FC10" bold="true" box="[899,976,925,948]" italics="true" pageId="13" pageNumber="456">Variola</emphasis>
Swainson
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(2 species)
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Type species: 
<taxonomicName id="4C570AB1E12BDF3EFC0038996828FC75" authority="Swainson" authorityName="Swainson" box="[995,1329,954,977]" family="Serranidae" genus="Variola" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="456" phylum="Chordata" rank="species" species="longipinna">
<emphasis id="B923AD20E12BDF3EFC00389969AAFC74" box="[995,1203,954,976]" italics="true" pageId="13" pageNumber="456">Variola longipinna</emphasis>
Swainson
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(
<emphasis id="B923AD20E12BDF3EFAAB389A6841FC74" bold="true" box="[1352,1368,953,976]" pageId="13" pageNumber="456">¼</emphasis>
<taxonomicName id="4C570AB1E12BDF3EFABB38986EBAFC49" authority="Forsskal" authorityName="Forsskal" authorityYear="1775" family="Percidae" genus="Perca" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="456" phylum="Chordata" rank="species" species="louti">
<emphasis id="B923AD20E12BDF3EFABB389868CBFC74" box="[1368,1490,954,976]" italics="true" pageId="13" pageNumber="456">Perca louti</emphasis>
Forsskål
</taxonomicName>
by monotypy).
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<paragraph id="8BE87132E12BDF3EFCD738D66969FBAE" blockId="13.[820,1490,925,1123]" box="[820,1136,1013,1035]" pageId="13" pageNumber="456">Type locality: Indian Ocean.</paragraph>
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<subSubSection id="C34D22B9E12BDF3EFCD73F316954FBC7" pageId="13" pageNumber="456" type="taxon_list">
<paragraph id="8BE87132E12BDF3EFCD73F316EE8FB8C" blockId="13.[820,1490,925,1123]" box="[820,1009,1042,1064]" pageId="13" pageNumber="456">Included species:</paragraph>
<paragraph id="8BE87132E12BDF3EFCAF3F13698CFBE1" blockId="13.[820,1490,925,1123]" box="[844,1173,1071,1093]" pageId="13" pageNumber="456">
<taxonomicName id="4C570AB1E12BDF3EFCAF3F13698CFBE1" authority="Baissac, 1953" authorityName="Baissac" authorityYear="1953" box="[844,1173,1071,1093]" family="Serranidae" genus="Variola" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="456" phylum="Chordata" rank="species" species="albimarginata">
<emphasis id="B923AD20E12BDF3EFCAF3F136EE3FBE1" box="[844,1018,1071,1093]" italics="true" pageId="13" pageNumber="456">V. albimarginata</emphasis>
Baissac, 1953
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</paragraph>
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<taxonomicName id="4C570AB1E12BDF3EFCAF3F6E6954FBC7" authority="(Forsskal, 1775)" baseAuthorityName="Forsskal" baseAuthorityYear="1775" box="[844,1101,1100,1123]" family="Percidae" genus="Perca" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="456" phylum="Chordata" rank="species" species="louti">
<emphasis id="B923AD20E12BDF3EFCAF3F6E6E81FBC6" box="[844,920,1100,1122]" italics="true" pageId="13" pageNumber="456">V. louti</emphasis>
(Forsskål, 1775)
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</paragraph>
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</treatment>
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