A revision of the spider genus Raveniola (Araneae, Nemesiidae). II. Species from Central Asia Author Zonstein, Sergei L. urn:lsid:zoobank.org:author:EADD3607-30FF-49AE-93F5-8410630469BE Steinhardt Museum of Natural History, Tel-Aviv University, 69978 Tel-Aviv, Israel znn@tauex.tau.ac.il text European Journal of Taxonomy 2024 2024-10-24 967 1 185 https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2699/12459 journal article 10.5852/ejt.2024.967.2699 2118-9773 13990819 C08B8027-50CC-417E-BCD4-5183B9FF6738 Raveniola caudata Zonstein, 2009 Figs 1 , 82 , 136 , 202 , 256–257 , 349 , 379–380 , 555 , 619–621, 748–749 Raveniola caudata Zonstein, 2009: 368 , figs 1, 3, 5–6, 8 ( ♂ ). Raveniola caudata – Mikhailov 2013: 12 . — Zonstein et al. 2018 : fig. 167 ( ♂ ). Diagnosis The species differs from Raveniola inopinata sp. nov. by having a considerably paler colour of ginger orange carapace and legs (which are dark sepia brown in the latter species; see Figs 1 and 2 ). Raveniola caudata can be distinguished from R . redikorzevi by its almost indistinct dorsal abdominal pattern. Males of R . caudata differ from males of the two other species of the group in having shorter laterodistal hair tufts on tarsi I–IV ( Fig. 257 cf. Figs 259 , 324 ) and by the shorter and less tapering proximal part of the embolus ( Figs 379–380 cf. Figs 381–384 ). Material examined Holotype TAJIKISTAN • ♂ ; Panj Karatau Mts, western slope of Mt Astana , 2.8 km WSW of summit; 37°22.8′ N , 69°12.8′ E ; 1020 m a.s.l. ; 24 Apr. 1991 ; S.V. Ovchinnikov leg.; SMNH . Additional material ( 1 ♂ , 1 juv. ) TAJIKISTAN • 1 juv. ; Aruktau Mts, surroundings of Ganjina ; 37°58′ N , 68°34′ E ; 700–800 m a.s.l. ; 16 Apr. 1968 ; V.F. Bahvalov leg.; SMNH • 1 ♂ (with both palps lost prior to collection); Vahsh Karatau Mts , 3 km NW of Mt Hojamaston ; 38°01.4′ N , 68°56.8′ E ; 940 m a.s.l. ; 21 Apr. 1989 ; S. Zonstein leg.; SMNH . Description Male ( holotype ) HABITUS . See Fig. 1. MEASUREMENTS . TBL 19.30, CL 7.13, CW 6.27, LL 0.57, LW 1.26, SL 3.52, SW 3.21. COLOUR . Carapace, palps and legs ginger orange; eye tubercle with central and two symmetrical lateral brownish-black spots surrounding AME and lateral eyes respectively; chelicerae cherry red; sternum, labium and maxillae light yellowish orange; metatarsi and tarsi gradually lighten toward apices; entire abdomen light yellowish grey, dorsally with almost indistinct darker pattern represented by few very weakly developed pairs of posteriorly-inclined greyish fasciae; spinnerets uniformly light yellowish grey. CEPHALOTHORAX . Carapace and chelicerae as shown in Fig. 82 . Clypeus and eye group as in Fig. 136 . Eye diameters and interdistances: AME 0.16(0.22), ALE 0.28, PLE 0.16, PME 0.12; AME–AME 0.16(0.10), ALE–AME 0.07(0.04), ALE–PLE 0.06, PLE–PME 0.04, PME–PME 0.43. Each cheliceral furrow with 10 promarginal teeth and 5 mesobasal denticles. MIT indiscernible. Sternum, labium and maxillae as shown in Fig. 202 . Maxillae with 52–57 cuspules each. LEGS . Tibia and metatarsus I as in Fig. 256 . Trichobothria: 2 rows of 9–12 each on tibiae, 13–15 on metatarsi, 15–18 on tarsi, 9–10 on cymbium. Scopula: distal ⅓ on metatarsi I–II, entire on tarsi I–II, widely divided by setae on tarsus III, absent on tarsus IV. Tarsi I–IV apically with very moderately dense lateral tufts of relatively short setae ( Fig 257 ). Paired claws on tarsi I–IV with 8–11 teeth on each margin. SPINATION . Palp: femur d4, pd3, rd2; patella p2; tibia d4, p3, r2, v4; cymbium d6(5). Leg I: femur d4, pd3, rd3; patella p2; tibia p3(1), pv2(1), r2+2M; metatarsus v1a. Leg II: femur d4, pd3; patella p2; tibia p3, v8(6); metatarsus p1; v6. Leg III: femur d4, pd3, rd3; patella p2, r1; tibia d2, p2, r2, v7(5); metatarsus d2, p3, r2, v10(9). Leg IV: femur d4, pd4(3), rd3; patella p2, r1; tibia d3(2), p3, r3(2), v7; metatarsus d3, p3, r3, v10(9). Tarsi I–IV aspinose. LEG MEASUREMENTS .

Femur	Patella	Tibia	Metatarsus	Tarsus	Total
Palp	4.23	2.27	3.46	–	1.35	11.31
Leg I	6.32	3.65	5.03	5.27	3.10	23.37
Leg II	6.22	3.23	4.98	5.02	3.03	22.48
Leg III	5.55	2.88	4.10	5.87	3.27	21.67
Leg IV	6.97	3.25	5.53	7.70	3.73	27.18

PALP . Tibia, cymbium and copulatory bulb as shown in Fig. 349 . Broadly tipped embolus with noticeably shortened basal part and with pronounced subapical keel ( Figs 379–380 ). SPINNERETS . See Fig. 555 . PMS: length 0.75; diameter 0.28. PLS: maximal diameter 0.65; length of basal, medial and apical segments 1.22, 0.85, 1.23; total length 3.30; apical segment elongate. Female Unknown. Ecology The species inhabits open shrubland and low forest biotopes with co-dominating Pistacia vera L., Cercis griffithii Boiss. , Acer spp. and Prunus spp. (see Figs 619–620 ). Males were found under rocks; the only juvenile specimen was found, according to the original label data, inside a gerbil’s burrow. The microbiotope situated directly in the type locality (shown in Fig. 621 ) is a pile of stones in the foreground, from where the holotype was collected in 1991. The corresponding photograph was taken in 2015. Distribution South Tajikistan , as shown in Figs 748–749 . In the original description, the distance between the ridge summit and the type locality was indicated incorrectly (see Zonstein 2009 ).