A Nomenclator of Croton (Euphorbiaceae) in Madagascar, the Comoros Archipelago, and the Mascarene Islands Author Berry, Paul E. Herbarium, Department of Ecology and Evolutionary Biology, University of Michigan, 3600 Varsity Drive, Ann Arbor, Michigan 48108, U. S. A. peberry@umich.edu Author Kainulainen, Kent Herbarium, Department of Ecology and Evolutionary Biology, University of Michigan, 3600 Varsity Drive, Ann Arbor, Michigan 48108, U. S. A. Author Ee, Benjamin W. van Department of Biology, Universidad de Puerto Rico, Recinto Universitario de Mayagueez, Mayagueez, PR 00680, Puerto Rico, U. S. A. text PhytoKeys 2017 2017-11-15 90 1 87 http://dx.doi.org/10.3897/phytokeys.90.20586 journal article http://dx.doi.org/10.3897/phytokeys.90.20586 1314-2003-90-1 80067D29FFFB7D34FF80E95D553F4254 1138341 19. Croton bemaranus Leandri, Ann. Mus. Colon. Marseille, ser . 5, 7(1): 69. 1939, as 'bemarana' Croton subaemulans var. tsingyensis Radcl.-Sm., Gen. Croton Madag. Comoro 120. 2016, syn. nov. Type. Madagascar. Prov. Mahajanga: Tsingy du Bemaraha (9e Reserve ), recu Feb -Apr 1933, J. Leandri 116 (holotype: K!; isotypes: P [P00133368!, P00133369!]). Type . Madagascar . Prov. Mahajanga : causse d'Ankara , Oct 1900 , H. Perrier de la Bathie 1153 ( lectotype , designated here: P [P00206491]!; isolectotypes: K [K001040396]!, P [P00389502]!) . Madagascar . Prov. Mahajanga : Bemara , partie Nord de l'Antsingy , entre Ambatomiloloha et Anjohinomby , 29 Nov 1932 , J. Leandri 656 ( syntypes : P [P00127512]!, P [P00389503]!) . Habit and distribution. Shrubs; western and northern Madagascar (Antsiranana, Mahajanga).