A new subgenus for “ Acalathus ” advena (LeConte, 1846) and the challenge of defining Calathina based on morphological characters (Coleoptera, Carabidae Sphodrini) Author Schmidt, Joachim Author Will, Kipling text Zootaxa 2020 2020-01-15 4722 4 326 338 journal article 24336 10.11646/zootaxa.4722.4.2 df6dd603-6441-4819-9023-f41c6511baef 1175-5326 3609581 558B1A4C-FB62-4D81-8571-C36C1A4AD061 Subgenus Bedelinus Ragusa, 1885 Third antennomere without pubescence. Mentum tooth simple. Pronotal laterobasal seta removed from side margin. Lateral bead of prosternal process present. Parascutellar seta present. Elytra glabrous, with two discal setae in the third interval; discal setae absent in the fifth and seventh intervals. Elytral microsculpture nearly isodiametric. Metepisternum elongated. Male protarsomeres dilated. Lateral grooves present on metatarsomeres I-IV. Apical gonocoxite short, robust, with two ensiform setae at external margin; sensory pit of apical gonocoxite well developed. Aedeagus left side up in repose; median lobe apical disc absent; apical hook of long paramere present; membranous lobe of short paramere present.