<document ID-DOI="http://dx.doi.org/10.3897/zookeys.144.62246" ID-GBIF-Dataset="3776e555-85f5-46ed-8ea1-98ca6a147ede" ID-PMC="PMC8222199" ID-Pensoft-Pub="1313-2970-1044-609" ID-Pensoft-UUID="C96ED07B505A52A8ADA354E1F70BB7C0" ID-PubMed="34183888" ID-ZooBank="66F01A49D32448A8AC2669BF3374894C" ModsDocID="1313-2970-1044-609" checkinTime="1623875354053" checkinUser="pensoft" docAuthor="Smith, Sarah M. &amp; Cognato, Anthony I." docDate="2021" docId="98A643694F81534BB5349E14AD4A1844" docLanguage="en" docName="ZooKeys 1044: 609-720" docOrigin="ZooKeys 1044" docPubDate="2021-06-16" docSource="http://dx.doi.org/10.3897/zookeys.144.62246" docTitle="Coptoborus ripley Smith &amp; Cognato 2021, sp. nov." docType="treatment" docUuid="C18AB0BB-FAED-4C67-A33D-A469E045C2DF" docUuidSource="ZooBank" docVersion="4" id="C96ED07B505A52A8ADA354E1F70BB7C0" lastPageNumber="609" masterDocId="C96ED07B505A52A8ADA354E1F70BB7C0" masterDocTitle="A revision of the Neotropical genus Coptoborus Hopkins (Coleoptera, Curculionidae, Scolytinae, Xyleborini)" masterLastPageNumber="720" masterPageNumber="609" pageNumber="609" updateTime="1668150492206" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the Neotropical genus Coptoborus Hopkins (Coleoptera, Curculionidae, Scolytinae, Xyleborini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Smith, Sarah M.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5173-3736</mods:nameIdentifier>
<mods:affiliation>Department of Entomology, Michigan State University, 288 Farm Lane, East Lansing, Michigan 48824, USA</mods:affiliation>
<mods:nameIdentifier type="email">camptocerus@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Cognato, Anthony I.</mods:namePart>
<mods:affiliation>Department of Entomology, Michigan State University, 288 Farm Lane, East Lansing, Michigan 48824, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-06-16</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1044</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>609</mods:start>
<mods:end>720</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.144.62246</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.144.62246</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1044-609</mods:identifier>
<mods:identifier type="ZooBank">66F01A49D32448A8AC2669BF3374894C</mods:identifier>
<mods:identifier type="Pensoft-UUID">C96ED07B505A52A8ADA354E1F70BB7C0</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="182753916" LSID="urn:lsid:zoobank.org:act:C18AB0BB-FAED-4C67-A33D-A469E045C2DF" httpUri="http://treatment.plazi.org/id/98A643694F81534BB5349E14AD4A1844" lastPageNumber="609" pageId="0" pageNumber="609">
<subSubSection pageId="0" pageNumber="609" type="nomenclature">
<paragraph pageId="0" pageNumber="609">
<taxonomicName LSID="http://zoobank.org/C18AB0BB-FAED-4C67-A33D-A469E045C2DF" authority="Smith &amp; Cognato, 2021" authorityName="Smith &amp; Cognato" authorityYear="2021" class="Insecta" family="Curculionidae" genus="Coptoborus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptoborus ripley" order="Coleoptera" pageId="0" pageNumber="609" phylum="Arthropoda" rank="species" species="ripley" status="sp. nov.">Coptoborus ripley</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="609">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Dorsal, lateral, frontal and declivital view of Coptoborus ricini, 2.3 - 2.5 mm (A-C, M), C. ripley holotype, 3.5 mm (D-F, N), C. sagitticauda holotype, 2.3 mm (G-I, O), C. sarahconnor holotype, 2.3 mm (J-L, P). All photographs by SMS." figureDoi="10.3897/zookeys.144.62246.figure14" httpUri="https://binary.pensoft.net/fig/555849" pageId="0" pageNumber="609">Figure 14D-F, N</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="609" type="type material">
<paragraph pageId="0" pageNumber="609">Type material.</paragraph>
<paragraph pageId="0" pageNumber="609">
<emphasis bold="true" pageId="0" pageNumber="609">
<emphasis italics="true" pageId="0" pageNumber="609">Holotype</emphasis>
,
</emphasis>
female, Ecuador: Napo Prov. [= Orellana Prov.], Res[erva]. Ethnica Waorani, 1 km S. Okone Gare Camp, Trans[ect]. Ent[omology]., 
<geoCoordinate degrees="00" direction="south" minutes="39" orientation="latitude" precision="15" seconds="10" value="-0.65277773">00°39'10&quot;S</geoCoordinate>
, 
<geoCoordinate degrees="076" direction="west" minutes="26" orientation="longitude" precision="925" value="-76.433334">076°26'W</geoCoordinate>
, 220 m, July 1996, T.L. Erwin et al. collectors, insecticidal fogging, terra firme forest, trans[ect] 3, sta[tion] 6, Erwin lot #1546 (ICB).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="609" type="diagnosis">
<paragraph pageId="0" pageNumber="609">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="609">
3.5 mm (n = 1), 2.69 
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide. This species is distinguished by the elytral apex attenuate and weakly emarginate, declivity distinctly sulcate along interstriae 2, declivital interstriae 2 unarmed and impunctate, interstriae 1 and 3 armed, declivital striae 1 and 2 parallel on declivital face and widely spaced, and large size.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="609" type="similar species">
<paragraph pageId="0" pageNumber="609">Similar species.</paragraph>
<paragraph pageId="0" pageNumber="609">
<taxonomicName family="Combretaceae" kingdom="Animalia" lsidName="C. busoror" order="Coleoptera" pageId="0" pageNumber="609" rank="species" species="busoror">
<emphasis italics="true" pageId="0" pageNumber="609">C. busoror</emphasis>
</taxonomicName>
, 
<taxonomicName family="Combretaceae" kingdom="Animalia" lsidName="C. leeloo" order="Coleoptera" pageId="0" pageNumber="609" rank="species" species="leeloo">
<emphasis italics="true" pageId="0" pageNumber="609">C. leeloo</emphasis>
</taxonomicName>
, 
<taxonomicName family="Combretaceae" kingdom="Animalia" lsidName="C. nudulus" order="Coleoptera" pageId="0" pageNumber="609" rank="species" species="nudulus">
<emphasis italics="true" pageId="0" pageNumber="609">C. nudulus</emphasis>
</taxonomicName>
, 
<taxonomicName family="Combretaceae" kingdom="Animalia" lsidName="C. ochromactonus" order="Coleoptera" pageId="0" pageNumber="609" rank="species" species="ochromactonus">
<emphasis italics="true" pageId="0" pageNumber="609">C. ochromactonus</emphasis>
</taxonomicName>
, 
<taxonomicName family="Combretaceae" kingdom="Animalia" lsidName="C. pilisoror" order="Coleoptera" pageId="0" pageNumber="609" rank="species" species="pilisoror">
<emphasis italics="true" pageId="0" pageNumber="609">C. pilisoror</emphasis>
</taxonomicName>
, 
<taxonomicName family="Combretaceae" kingdom="Animalia" lsidName="C. sororcula" order="Coleoptera" pageId="0" pageNumber="609" rank="species" species="sororcula">
<emphasis italics="true" pageId="0" pageNumber="609">C. sororcula</emphasis>
</taxonomicName>
, 
<taxonomicName family="Combretaceae" kingdom="Animalia" lsidName="C. spicatus" order="Coleoptera" pageId="0" pageNumber="609" rank="species" species="spicatus">
<emphasis italics="true" pageId="0" pageNumber="609">C. spicatus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="609" type="description">
<paragraph pageId="0" pageNumber="609">Description</paragraph>
<paragraph pageId="0" pageNumber="609">
<emphasis bold="true" pageId="0" pageNumber="609">
(female). 
<emphasis italics="true" pageId="0" pageNumber="609">Holotype</emphasis>
</emphasis>
3.5 mm, 2.69 
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide. Body dark brown, antennae and legs lighter. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="609">Head</emphasis>
: epistoma smooth. Frons subshiny, finely punctate, setose; each puncture bearing a long, erect hair-like seta. Eyes broadly and moderately emarginate. Submentum large, triangular, slightly impressed. Antennal scape regularly thick, as long as club. Pedicel shorter than funicle. Club circular, flat, type 3; segment 1 corneous, transverse on anterior face, occupying basal ~1/3; segment 2 narrow, transverse, corneous; segments 1 and 2 present on posterior face. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="609">Pronotum</emphasis>
: 1.2 
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide. In dorsal view long and rounded frontally, type 7, sides parallel in basal 3/5, rounded anteriorly; anterior margin with two projecting serrations. In lateral view elongate, disc longer than anterior slope, type 7, summit prominent, on anterior 3/5. Anterior slope with densely spaced, broad fine asperities, becoming lower and more strongly transverse towards summit. Disc reticulate, dull with sparse, minute punctures, some longer hair-like setae at margins. Lateral margins obliquely costate. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="609">Elytra</emphasis>
: 1.5 
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide, 1.3 
<normalizedToken originalValue="×">x</normalizedToken>
as long as pronotum. Scutellum minute. Elytra attenuate, parallel-sided in basal 70%, then acutely tapered to apex, apex strongly emarginate. Disc smooth, shiny; strial punctures moderate, shallow, glabrous; interstriae flat, sparsely, minutely punctate, unarmed, each puncture bearing a long semi-erect hair-like seta. Declivity gradual, smooth, shiny, appearing bisulcate, occupying apical 2/5 of elytra; striae not impressed, striae 1 and 2 parallel, strial punctures much larger and shallower than those of disc; interstriae impunctate, interstriae 2 strongly sulcate, unarmed; interstriae 1 and 3 strongly costate with five and three granules respectively, each granule bearing a long, erect hair-like seta. Posterolateral margin with interstriae 3 and 9 joining, forming a granulate acute carina and continuing submarginally to apex. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="609">Legs</emphasis>
: protibiae obliquely triangular, broadest at apical 1/4; apical 1/2 of outer margin with six large, socketed denticles, their length longer than basal width. Meso- and metatibiae flattened; outer margin evenly rounded with seven large, socketed denticles.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="609" type="etymology">
<paragraph pageId="0" pageNumber="609">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="609">
Portrayed by Sigourney Weaver, Ellen Ripley is the heroine in the movies 
<normalizedToken originalValue="‘Alien’">'Alien'</normalizedToken>
(1979), 
<normalizedToken originalValue="‘Aliens’">'Aliens'</normalizedToken>
(1986), 'Alien 3' (1992), and 'Alien: 
<normalizedToken originalValue="Resurrection’">Resurrection'</normalizedToken>
(1997). This species is glabrous and reminiscent of 
<normalizedToken originalValue="Ripley’s">Ripley's</normalizedToken>
shaved head in 'Alien 3'. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="609" type="distribution">
<paragraph pageId="0" pageNumber="609">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="609">Ecuador (Orellana).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="609" type="biology_ecology">
<paragraph pageId="0" pageNumber="609">Biology.</paragraph>
<paragraph pageId="0" pageNumber="609">The holotype was collected by canopy fogging.</paragraph>
</subSubSection>
</treatment>
</document>