New southern African species and a revalidation in the dung beetle genus Gyronotus van Lansberge, 1874 (Scarabaeidae: Scarabaeinae) with an updated key Author Deschodt, Christian M. Author Davis, Adrian L. V. text Zootaxa 2019 2019-06-28 4624 2 275 282 journal article 26368 10.11646/zootaxa.4624.2.10 7fc3c435-d3ed-45db-bd7d-48398d831f01 1175-5326 3259222 B54840C3-4AFE-475E-8B99-7FA8FD799645 Revalidation of Gyronotus marginatus Péringuey, 1888 ( Figs. 1E , 3 ) Although Scholtz & Howden (1987: 82) did not see the type of G. marginatus , they stated that “ Gyronotus marginatus Péringuey is without doubt a synonym of G. pumilus . Although the type is missing from the Péringuey collection it is clear from the description and the type locality that the two names refer to one species. Consequently we synonymize G. marginatus with G. pumilus ”. The type of Gyronotus marginatus has subsequently been located in the Iziko South African Museum (ISAM). It is clearly larger than any specimens of Gyronotus pumilus , as are the paratype specimens (examined by ALVD) that are housed in BMNH. Consequently, we revalidate Gyronotus marginatus from the junior subjective synonymy with Gyronotus pumilus and re-elevate it to species rank. Diagnosis. Gyronotus marginatus is clearly much bigger than Gyronotus pumilus . Also, the elytra of G. marginatus are shagreened and fairly even while the elytra of G. pumilus are shagreened and somewhat rugose.