A revision of the Cossulinae of Costa Rica and cladistic analysis of the world species (Lepidoptera: Cossidae) Author Davis, Steve R. Author Gentili-Poole, Patricia Author Mitter, Charles text Zoological Journal of the Linnean Society 2008 2008-10-31 154 2 222 277 https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00406.x journal article 10.1111/j.1096-3642.2008.00406.x 0024-4082 5446371 SIMPLICIVALVA AMPLIOPHILOBIA SP. NOV. ( FIGS 31 , 67, 68 , 96 , 139 ) Male ( Fig. 31 ): Forewing length: 18.5– 20 mm . Head: Antennae cream; labial palpi fuscous dorsally, cream and light brown ventrally; frons cream, irrorated with fuscous and black; vertex same as frons. Thorax: Patagia fuscous to black, irrorated with white; tegula and dorsum cream, irrorated with fuscous and black; dorsum with a rectangular fuscous-black patch along midline; pleura cream, with white irrorated with fuscous-black along base of forewing; venter mostly cream; a dark brown to fuscous patch below labial palps, ventrally. Fore and mid legs with femur and tibia cream, irrorated with brown; hind legs with femur and tibia cream; tarsi brown, banded with cream distally on each tarsomere. Forewing cream dorsally, with a few steely-grey striae interspersed throughout wing; costal margin cream and brown banded; three large steely-grey spots immediately before half length of wing, the first in middle of discal cell, the second between CuP and 1A+2A, and the third at inner margin, below 1A+2A; terminal third of wing cream, interspersed with rusty orange; immediately before termen, a large steelygrey patch, and before this patch, two large fuscous spots; fringe steely-grey and cream; forewing different shades of brown and light brown ventrally, interspersed with cream; costal margin cream and brown banded; all spots visible dorsally are also present ventrally, but fainter in appearance. Hindwing intermixed with brown and cream dorsally; hindwing mostly cream ventrally, mixed with brown, and light brown patches along termen at vein endings; fringe mostly cream, slightly banded steel-grey. Abdomen: Cream and light brown dorsally, with A1–A2 slightly darker brown; cream and light brown ventrally. Male genitalia ( Figs 67, 68 , 96 ): Valva oblong and rounded terminally, with terminal end slightly upcurved; length of valva approximately equal to length of genital capsule; gnathos–uncus expanse approximately 0.7¥ length of genital capsule; sacculus with a protruding ridge that is somewhat fattened, but nothing more than a ridge; juxta process short, flat and wide; gnathos arms elongate and slender, broadly curving as viewed laterally; gnathos bridge elongate and narrow as viewed posterio-ventrally, blunt and lobed terminally; uncus elongate and narrow, becoming bulbous terminally, with apex truncate and very shallowly bifurcate. Aedoeagus becoming slightly wider towards rostellum, as viewed laterally; rostellum elongate and bluntly rounded teminally; vesica with a dense covering of minute, setiform spines. Female: Unknown. Distribution: Known only from provinces of Puntarenas and San Jose , Costa Rica . Holotype : ♂ ; COSTA RICA : SAN JOSÉ : Estación Zurqui ( El Tunel ), Parque Nacional Braulio Carrillo , 1500 m : 9–11.vi.1986 , A. Chacón ( INBio ). Paratype : COSTA RICA : PUNTARENAS : Finca Cafrosa , Estación Las Mellizas , Parque Nacional Amistad , 1300 m : 1 ♂ , vi.–vii.1990 , J. C. Saborio , slide USNM 85636 ( USNM ) . Host: Unknown. Flight period: June and July. Etymology: Derived from the Latin ‘amplius’ meaning ‘larger’. Simplicivalva ampliophilobia refers to the larger size of this species as compared with the very similar, smaller species S. philobia . Discussion: This species most closely resembles two species not known to occur in Costa Rica , S. eberti and S. philobia from Brazil . Simplicivalva ampliophilobia differs from both species by being larger in size and possessing two large, nearly contiguous, fuscous spots near the termen of the forewing. In both S. eberti and S. philobia these spots are fused.